Wheeler, W. M. 1903b. Ethological observations on an American ant (Leptothorax Emersoni Wheeler). [part]. Journal für Psychologie und Neurologie 2:31-47.

Wheeler, W. M. 1903c. A decad of Texan Formicidae. Psyche (Cambridge) 10:93-111. PDF

Notes: Stamp date in MCZ library: 1906.06.30.

Wheeler, W. M. 1903d. A revision of the North American ants of the genus Leptothorax Mayr. Proceedings of the Academy of Natural Sciences of Philadelphia 55:215-260. PDF

Wheeler, W. M. 1903e. Ethological observations on an American ant (Leptothorax Emersoni Wheeler). [concl.]. Journal für Psychologie und Neurologie 2:64-78.

Wheeler, W. M. 1903f. The North American ants of the genus Stenamma sensu stricto. Psyche (Cambridge) 10:164-168. PDF

Notes: Stamp date in MCZ library: 1903.08.22.

Wheeler, W. M. 1903g. Extraordinary females in three species of Formica, with remarks on mutation in the Formicidae. Bulletin of the American Museum of Natural History 19:639-651. PDF

Wheeler, W. M. 1903h. Some new gynandromorphous ants, with a review of the previously recorded cases. Bulletin of the American Museum of Natural History 19:653-683. PDF

Wheeler, W. M. 1903i. The origin of female and worker ants from the eggs of parthenogenetic workers. Science (New York) (n.s.)18:830-833.

Wheeler, W. M. 1903j. Some notes on the habits of Ceraphachys augustae. Psyche (Cambridge) 10:205-209. PDF

Notes: Stamp date in MCZ library: 1904.01.04.

Wheeler, W. M. 1904a. Three new genera of inquiline ants from Utah and Colorado. Bulletin of the American Museum of Natural History 20:1-17. PDF

Wheeler, W. M. 1904b. Dr. Castle and the Dzierzon theory. Science (New York) (n.s.)19:587-591. PDF

Wheeler, W. M. 1904c. The American ants of the subgenus Colobopsis. Bulletin of the American Museum of Natural History 20:139-158. PDF

Wheeler, W. M. 1904d. A crustacean-eating ant (Leptogenys elongata Buckley). Biological Bulletin (Woods Hole) 6:251-259. PDF

Wheeler, W. M. 1904e. Ants from Catalina Island, California. Bulletin of the American Museum of Natural History 20:269-271. PDF

Wheeler, W. M. 1904f. The ants of North Carolina. Bulletin of the American Museum of Natural History 20:299-306. PDF

Wheeler, W. M. 1904g. On the pupation of ants and the feasibility of establishing the Guatemalan Kelep, or Cotton-Weevil Ant, in the United States. Science (New York) (n.s.)20:437-440. PDF

Wheeler, W. M. 1904h. Social parasitism among ants. American Museum Journal 4:74-75.

Wheeler, W. M. 1904i. A new type of social parasitism among ants. Bulletin of the American Museum of Natural History 20:347-375. PDF

Wheeler, W. M. 1904j. Some further comments on the Guatemalan boll weevil ant. Science (New York) (n.s.)20:766-768. PDF

Wheeler, W. M. 1905a. An interpretation of the slave-making instincts in ants. Bulletin of the American Museum of Natural History 21:1-16. PDF

Wheeler, W. M. 1905b. Ants from Catalina Island, Cal. Bulletin of the Southern California Academy of Sciences 4:60-62.

Notes: Reprint of Wheeler (1904e) with some additions and emendations.

Wheeler, W. M. 1905c. The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History 21:79-135. PDF

Wheeler, W. M. 1905d. New species of Formica. Bulletin of the American Museum of Natural History 21:267-274. PDF

Wheeler, W. M. 1905e. How the queens of the parasitic and slave-making ants establish their colonies. American Museum Journal 5:144-148. PDF

Wheeler, W. M. 1905f. Some remarks on temporary social parasitism and the phylogeny of slavery among ants. Biologisches Centralblatt 25:637-644. PDF

Wheeler, W. M. 1905g. The North American ants of the genus Dolichoderus. Bulletin of the American Museum of Natural History 21:305-319. PDF

Wheeler, W. M. 1905h. The North American ants of the genus Liometopum. Bulletin of the American Museum of Natural History 21:321-333. PDF

Wheeler, W. M. 1905i. Dr. O. F. Cook's "Social organization and breeding habits of the cotton-protecting kelep of Guatemala.". Science (New York) (n.s.)21:706-710. PDF

Wheeler, W. M. 1905j. An annotated list of the ants of New Jersey. Bulletin of the American Museum of Natural History 21:371-403. PDF

Wheeler, W. M. 1905k. Worker ants with vestiges of wings. Bulletin of the American Museum of Natural History 21:405-408.