Cardenas, C. R.; Luo, A. R.; Jones, T. H.; Schultz, T. R.; Adams, R. M. M. 2021. Using an integrative taxonomic approach to delimit a sibling species, Mycetomoellerius mikromelanos sp. nov. (Formicidae: Attini: Attina). Proceedings of the National Academy of Sciences of the United States of America 9:e11622. 10.7717/peerj.11622 PDF

Aswaj, P.; Anoop, K.; Priyadarsanan, D. R. 2021. Description of two new species of ants of the genus Myrmecina Curtis, 1829 (Hymenoptera: Formicidae: Myrmicinae) from the Eastern Himalayas. Zootaxa 4990:160-171. 10.11646/zootaxa.4990.1.10 PDF

Akbar, S. A.; Bharti, H.; Kanturski, M.; Wachkoo, A. A. 2021. First record of the myrmicine ant genus Syllophopsis Santschi, 1915 (Hymenoptera: Formicidae) from India with description of a new species. Zootaxa 4985:403-413. 10.11646/ZOOTAXA.4985.3.7 PDF

Wang, C.; Lin, C.-C.; Billen, J. 2021. Morphology of the novel ventral scape gland in the ant genus Strumigenys. Arthropod Structure & Development 63:7 pp. 101063. [online early] 10.1016/j.asd.2021.101063 PDF

General, D. E. M.; Buenavente, P. A. C.; Peeters, C. 2020. The twists and turns of biological research: first record of an ergatoid queen in the ant genus Pheidole Westwood, 1839, and implications for dispersal and life history. Asian Myrmecology 13:10 pp. e013002. [online early] 10.20362/am.013002 PDF

Satria, R.; Jannatan, R. 2021. Dealate queens of the ant genus Eurhopalothrix Brown et Kempf, 1961 (Hymenoptera: Formicidae: Myrmicinae) from Sumatra. Far Eastern Entomologist 430:11-16. 10.25221/fee.430.3 PDF

Notes: Mark queen descriptions

Booher, D. B.; Hoenle, P. O. 2021. A new species group of Strumigenys (Hymenoptera, Formicidae) from Ecuador, with a description of its mandible morphology. ZooKeys 1036:1-19. 10.3897/zookeys.1036.62034 PDF

Seifert, B. 2021. Surviving the winter: Tetramorium sibiricum n. sp., a new Central Siberian ant species (Hymenoptera: Formicidae). Osmia 9:15-24. 10.47446/OSMIA9.3 PDF

Radchenko, A. G.; Perkovsky, E. E. 2021. [Formica ribbeckei sp. nov. attributed to Radchenko & Perkovsky]. Pp. 240-241 in: Radchenko, A. G.; Perkovsky, E. E.; Vasilenko, D. V. 2021. Formica species (Hymenoptera, Formicidae, Formicinae) in late Eocene Rovno amber. Journal of Hymenoptera Research 82:237-251. 10.3897/jhr.82.64599 PDF

Radchenko, A. G.; Perkovsky, E. E.; Vasilenko, D. V. 2021. Formica species (Hymenoptera, Formicidae, Formicinae) in late Eocene Rovno amber. Journal of Hymenoptera Research 82:237-251. 10.3897/jhr.82.64599 PDF

Seifert, B. 2021. A taxonomic revision of the Palaearctic members of the Formica rufa group (Hymenoptera: Formicidae) – the famous mound-building red wood ants. Myrmecological News 31:133-179. 10.25849/myrmecol.news_031:133 PDF

Nsengimana, V.; Vanderheyden, A.; Gombeer, S.; Smitz, N.; Meganck, K.; De Meyer, M.; Backeljau, T.; Fisher, B. L.; Dekoninck, W. 2021. First record of the ant Pheidole megatron Fischer and Fisher, 2013 (Hymenoptera: Formicidae) from Rwanda. African Zoology 1:1-5. [online early] 10.1080/15627020.2021.1901605 PDF

Escárraga, M. E.; Lattke, J. E.; Pie, M. R.; Guerrero, R. J. 2021. Morphological and genetic evidence supports the separation of two Tapinoma ants (Formicidae, Dolichoderinae) from the Atlantic Forest biome. ZooKeys 1033:35-62. 10.3897/zookeys.1033.59880 PDF

Ward, P. S.; Boudinot, B. E. 2021. Grappling with homoplasy: taxonomic refinements and reassignments in the ant genera Camponotus and Colobopsis (Hymenoptera: Formicidae). Arthropod Systematics and Phylogeny 79:37-56. 10.3897/asp.79.e66978 PDF

Satria, R.; Herwina, H. 2020. New distribution records of ant species (Hymenoptera: Formicidae) to the fauna of Sumatra Island, Indonesia. Advances in Biological Sciences Research 10:82-84. PDF

Wang, W. Y.; Jaitrong, W. 2021. [Untitled. New species in Rhopalomastix attributed to Wang, W.Y. & Jaitrong, W.]. Pp. 123-155 in: Wang, W. Y.; Yong, G. W. J.; Jaitrong, W. 2021. Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species. European Journal of Taxonomy 739:117-157. PDF

Wang, W. Y.; Yong, G. W. J.; Jaitrong, W. 2021. Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species. European Journal of Taxonomy 739:117-157. 10.5852/ejt.2021.739.1271 PDF

Schifani, E.; Costa, S.; Mei, M.; Alicata, A. 2021. A new species for the Italian fauna: Aphaenogaster strioloides, not A. crocea, inhabits Pantelleria Island (Hymenoptera: Formicidae). Fragmenta Entomologica 53 (1):21-24. 10.13133/2284-4880/482 PDF

Varela-Hernández, F.; Riquelme, F. 2021. A new ant species of the genus Pheidole Westwood, 1839 from Miocene Mexican amber. Southwestern Entomologist 46 (1):75-82. 10.3958/059.046.0107 PDF

Radchenko, A. G.; Perkovsky, E. E. 2021. Wheeler's dilemma revisited: first Oecophylla–Lasius syninclusion and other ants syninclusions in the Bitterfeld amber (late Eocene). Invertebrate Zoology 18 (1):47-65. 10.15298/invert-zool.18.1.05 PDF

Notes: Age of Bitterfeld amber remains equivocal between late Eocene and Oligocene

Fernandes, I. O.; Larabee, F. J.; Oliveira, M. L.; Delabie, J. H. C.; Schultz, T. R. 2021. A global phylogenetic analysis of trap-jaw ants, Anochetus Mayr and Odontomachus Latreille (Hymenoptera: Formicidae: Ponerinae). Systematic Entomology (online early):1-19. [online early] 10.1111/syen.12483 PDF

Richter, A.; Hita Garcia, F.; Keller, R. A.; Billen, J.; Katzke, J.; Boudinot, B. E.; Economo, E. P.; Beutel, R. G. 2021. The head anatomy of Protanilla lini (Hymenoptera: Formicidae: Leptanillinae), with a hypothesis of their mandibular movement. Myrmecological News 31:85-114. 10.25849/myrmecol.news_031:85 PDF

Notes: Head anatomy, Protanilla

Al-Keridis, L. A.; Gaber, N. M.; Aldawood, A. S.; Sharaf, M. R. 2021. Description of the queen of the Arabian endemic dolichoderine species Tapinoma wilsoni Sharaf & Aldawood, 2012 (Hymenoptera: Formicidae) with the first illustrated key to queens of the Arabian Tapinoma Foerster, 1850. African Entomology 29 (1):233-237. 10.4001/003.029.0233 PDF

Notes: Tapinoma queen key to species

Billen, J.; Khalife, A.; Ito, F.; Anh, N.D.; Esteves, F. A. 2021. The basitarsal sulcus gland, a novel exocrine structure in ants. Arthropod Structure & Development 61:1-10. [online early] 10.1016/j.asd.2021.101041 PDF

Notes: Basitarsal sulcus gland anatomy

Perfilieva, K. S. 2021. Distribution and differentiation of fossil Oecophylla (Hymenoptera: Formicidae) species by wings imprints. Paleontological Journal 55:76-89. 10.1134/S003103012101010X PDF

Longino, J. T.; Branstetter, M. G. 2021. Integrating UCE phylogenomics with traditional taxonomy reveals a trove of New World Syscia species (Formicidae: Dorylinae). Insect Systematics and Diversity 5(2):1-51. 10.1093/isd/ixab001 PDF

Day, M. C.; Fitton, M. G. 1978. Re-curation of the Linnaean Hymenoptera (Insecta), with a reassessment of the taxonomic importance of the collection. Biological Journal of the Linnean Society 10 (2):181-198. 10.1111/j.1095-8312.1978.tb00012.x PDF

Csősz, S.; Fisher, B. L. 2021. New species of Aphaenogaster attributed to Csősz & Fisher]. Pp. 15-29 in: Csősz, S.; Loss, A. C.; Fisher, B. L. 2021. Taxonomic revision of the Malagasy Aphaenogaster swammerdami group (Hymenoptera: Formicidae). PeerJ 9:e10900:1-34. PDF

Csősz, S.; Loss, A. C.; Fisher, B. L. 2021. Taxonomic revision of the Malagasy Aphaenogaster swammerdami group (Hymenoptera: Formicidae). PeerJ 9:e10900:1-34. DOI 10.7717/peerj.10900 PDF

Tinaut, A.; Reyes-López, J. L. 2020. Descripción de una nueva especie para la península ibérica: Temnothorax alfacarensis n. sp. (Hymenoptera, Formicidae). Boletín de la Asociación Española de Entomologia 44:359-378. PDF