Schoeters, E. 1990. Morphology of the glandular system in the leaf-cutting ants Atta sexdens sexdens (L., 1758) and Atta sexdens rubropilosa Forel, 1908 (Hymenoptera: Formicidae). [Abstract.]. Belgian Journal of Zoology 120(Suppl. 1):52-53.

Schoeters, E.; Billen, J. P. J. 1990. Morphology of the venom gland in relation to worker size in leaf-cutting ants (Formicidae, Attini). Actes des Colloques Insectes Sociaux 6:249-252.

Schoeters, E.; Billen, J. P. J. 1991. Morphologie des glandes pro- et postpharyngiennes chez Atta sexdens (Hymenoptera: Formicidae). Actes des Colloques Insectes Sociaux 7:153-160.

Schoeters, E.; Billen, J. P. J. 1992. Morphological and ultrastructural study of the metapleural gland in Diacamma (Hymenoptera, Formicidae). Pp. 239-247 in: Billen, J. P. J. (ed.) 1992. Biology and evolution of social insects. Leuven: Leuven University Press, ix + 390 pp.

Schoeters, E.; Billen, J. P. J. 1993a. Anatomy and fine structure of the metapleural gland in Atta (Hymenoptera, Formicidae). Belgian Journal of Zoology 123:67-75.

Schoeters, E.; Billen, J. P. J. 1993b. Glandes coxales de Pachycondyla obscuricornis (Formicidae, Ponerinae). Actes des Colloques Insectes Sociaux 8:183-186.

Billen, J. P. J.; Schoeters, E. 1994. Morphology and ultrastructure of the mandibular gland in Formica L. ants (Hymenoptera, Formicidae). Memorabilia Zoologica 48:9-16.

Oldham, N. J.; Morgan, E. D.; Gobin, B.; Schoeters, E.; Billen, J. P. J. 1994. Volatile secretions of Old World army ant Aenictus rotundatus and chemotaxonomic implications of army ant Dufour gland chemistry. Journal of Chemical Ecology 20:3297-3305.

Schoeters, E.; Billen, J. P. J. 1994. Nouvelles données structurales sur la glande à venin chez Atta sexdens rubropilosa et Myrmica sabuleti (Hymenoptera, Formicidae). Actes des Colloques Insectes Sociaux 9:67-73.

Schoeters, E.; Billen, J. P. J. 1994a. The intramandibular gland, a novel exocrine structure in ants (Insecta, Hymenoptera). Zoomorphology (Berlin) 114:125-131.

Schoeters, E.; Billen, J. P. J. 1994b. The venom gland in formicine ants: an ontogenetic and ultrastructural study. P. 218 in: Lenoir, A.; Arnold, G.; Lepage, M. (eds.) 1994. Les insectes sociaux. 12ème congrès de l'Union Internationale pour l'Étude des Insectes Sociaux. Paris, Sorbonne, 21-27 août 1994. Paris: Université Paris Nord, xxiv + 583 pp.

Schoeters, E.; Billen, J. P. J. 1995. Morphology and ultrastructure of the convoluted gland in the ant Dinoponera australis (Hymenoptera: Formicidae). International Journal of Insect Morphology and Embryology 24:323-332.

Nascimento, R. R. do; Schoeters, E.; Morgan, E. D.; Billen, J.; Stradling, D. J. 1996. Chemistry of metapleural gland secretions of three attine ants, Atta sexdens rubropilosa, Atta cephalotes, and Acromyrmex octospinosus (Hymenoptera: Formicidae). Journal of Chemical Ecology 22:987-1000.

Wenseleers, T.; Schoeters, E.; Billen, J.; Hefetz, A. 1996. Morphologie et ultrastructure de la glande cloacale chez Cataglyphis niger. Actes des Colloques Insectes Sociaux 10:189-194.

Schoeters, E.; Billen, J. P. J. 1996a. The post-pharyngeal gland in Dinoponera ants (Hymenoptera: Formicidae): unusual morphology and changes during the secretory process. International Journal of Insect Morphology and Embryology 25:443-447.

Schoeters, E.; Billen, J. P. J. 1996b. The control apparatus of the venom gland in formicinae ants (Hymenoptera: Formicidae). Netherlands Journal of Zoology 46:281-289.

Wenseleers, T.; Schoeters, E.; Billen, J.; Wehner, R. 1998. Distribution and comparative morphology of the cloacal gland in ants (Hymenoptera: Formicidae). International Journal of Insect Morphology and Embryology 27:121-128.

Schoeters, E.; Ito, F.; Miyata, H.; Billen, J. P. J. 1999. Aberrant venom glands in Amblyoponini (Formicidae, Ponerinae): morphology, ultrastructure and histochemistry. Acta Zoologica (Stockholm) 80:3-9.

Schoeters, E.; Billen, J. P. J. 1999. Les organes neurohemaux perisympathiques des fourmis et differences entre les castes femelles. Actes des Colloques Insectes Sociaux 12:139-142. PDF

Billen, J.; Ito, F.; Tsuji, K.; Schoeters, E.; Maile, R.; Morgan, E. D. 2000. Structure and chemistry of the Dufour gland in Pristomyrmex ants (Hymenoptera, Formicidae). Acta Zoologica (Stockholm) 81:159-166.

Lommelen, E.; Schoeters, E.; Billen, J. 2003. Development of the labial gland of the ponerine ant Pachycondyla obscuricornis (Hymenoptera, Formicidae) during the pupal stage. Arthropod Structure and Development 32:209-217. 10.1016/S1467-8039(03)00052-5 PDF