References by Billen, J. P. J.
Wang, C.; Lin, C.-C.; Billen, J. 2021. Morphology of the novel ventral scape gland in the ant genus Strumigenys. Arthropod Structure & Development 63:7 pp. 101063. [online early] 10.1016/j.asd.2021.101063 PDF
Lieberman, Z. E.; Billen, J. P. J.; van de Kamp, T.; Boudinot, B. E. 2022. The ant abdomen: The skeletomuscular and soft tissue anatomy of Amblyopone australis workers (Hymenoptera: Formicidae). Journal of Morphology 283:693-770. 10.1002/jmor.21471 PDF

Notes: June 2022 issue. Published online 3 April 2022.

Billen, J. P. J.; Vanelderen, N.; Ito, F. 2022. Survey of the exocrine glands in workers of the ant Leptanilla clypeata. Insectes Sociaux -:1-13. 10.1007/s00040-022-00861 PDF
Ito, F.; Hashim, R.; Mizuno, R.; Billen, J. 2022. Notes on the biology of Protanilla sp. (Hymenoptera, Formicidae) collected in Ulu Gombak, Peninsular Malaysia. Insectes Sociaux 69:13-18. 10.1007/s00040-021-00839-z PDF

Notes: Published "online early" 27 Oct 2021, without volume number or final pagination.

Richter, A.; Boudinot, B. E.; Hita Garcia, F.; Billen, J.; Economo, E. P.; Beutel, R. G. 2023. Wonderfully weird: the head anatomy of the armadillo ant, Tatuidris tatusia (Hymenoptera: Formicidae: Agroecomyrmecinae), with evolutionary implications. Myrmecological News 33:35-75. 10.25849/myrmecol.news_033:035 PDF
Wang, C.; Chung, F.-Y.; Lin, C.-C.; Gibson, J. C.; McGuire, S.; Suarez, A.V.; Billen, J. 2023. The spongiform tissue in Strumigenys ants contains exocrine glands. Arthropod Structure and Development 73 (2023) 101246:10 pp. [online early] 10.1016/j.asd.2023.101246 PDF
Billen, J. P. J.; Wang, C. 2025. Intramandibular glands in ants. Myrmecological News 35:125-144. 10.25849/myrmecol.news_035:125 PDF
Protonyms originally described by author

Found no protonyms described by this author.

Statistics
Published between 1982–2025
Taxon descriptions between –
No. of references 127
No. of described taxa (including combinations) –
No. of described taxa (valid, including combinations) –
No. of described taxa (protonyms only) –
Names belonging to author
Author name No. of references
Billen, J. P. J. 95
Billen, J. 32