Billen, J. P. J.; Bauweleers, E.; Hashim, R.; Ito, F. 2013. Survey of the exocrine system in Protanilla wallacei (Hymenoptera, Formicidae). Arthropod Structure and Development 42:173-183. PDF

Ito, F.; Yamane, S. 2014. Reproduction by ergatoid queens in the myrmicine ant Monomorium brocha (Bolton) (Hymenoptera: Formicidae) in West Java, Indonesia, with a description of the male. Asian Myrmecology 6:105-113. PDF

Terayama, M.; Ito, F. 2014. A sibling species of Odontomachus rixosus Smith, 1857 (Hymenoptera: Formicidae) from Malaysia. Japanese Journal of Systematic Entomology 20 (2):181-183. PDF

Eguchi, K.; Mizuno R.; Ito, F.; Satria, R.; An, D. V.; Viet, B. T.; Luong, P. T. H. 2016. First discovery of subdichthadiigyne in Yunodorylus Xu, 2000 (Formicidae: Dorylinae). Revue Suisse de Zoologie 123 (2):307-314. 10.5281/zenodo.155307 PDF

Billen, J.; Ito, F. 2018. Novel thoracic glands in the ant Myopias hollandi. Arthropod Structure and Development 47:229-237. 10.1016/j.asd.2018.04.005 PDF

Yamada, M.; Ito, F.; Hashim, R.; Eguchi, K. 2018. Queen polymorphism in Acanthomyrmex careoscrobis Moffett, 1986 in Peninsular Malaysia (Hymenoptera: Formicidae: Myrmicinae), with descriptions of hitherto unknown female castes and males. Asian Myrmecology 10:19 pp. e010009. 10.20362/am.010009 PDF

Ito, F.; Nguyen, D. A.; Eguchi, K.; Toyota, A.; Billen, J. 2020. Colony composition, queen behavior, specialized predation on millipedes, and exocrine glands in the ponerine ant Myopias conicara Xu, 1998 (Hymenoptera: Formicidae). Entomologist's Record and Journal of Variation -:1-10. [online early] 10.1111/ens.12406 PDF

Ito, F.; Yamane, S. 2020. Behavior of the queen of Leptanilla clypeata Yamane et Ito collected in the Bogor Botanical Gardens, West Java, Indonesia (Hymenoptera; Formicidae), with a note on colony composition and a description of the ergatoid queen. Asian Myrmecology 12:7 pp. e012004. 10.20362/am.012004 PDF

Notes: Leptanilla clypeata ergatoid queen description

Billen, J.; Khalife, A.; Ito, F.; Anh, N.D.; Esteves, F. A. 2021. The basitarsal sulcus gland, a novel exocrine structure in ants. Arthropod Structure & Development 61: Article 101041:1-10. 10.1016/j.asd.2021.101041 PDF

Notes: Basitarsal sulcus gland anatomy

Ito, F.; Hashim, R.; Mizuno, R.; Billen, J. 2022. Notes on the biology of Protanilla sp. (Hymenoptera, Formicidae) collected in Ulu Gombak, Peninsular Malaysia. Insectes Sociaux 69:13-18. 10.1007/s00040-021-00839-z PDF

Notes: Published "online early" 27 Oct 2021, without volume number or final pagination.

Yamane, S.; Hosoishi, S.; Ito, F. 2022. Japanese Tetramorium queens: identification key and species diagnoses (Hymenoptera, Formicidae, Myrmicinae). ZooKeys 1084:43-64. 10.3897/zookeys.1084.69767 PDF

Billen, J. P. J.; Vanelderen, N.; Ito, F. 2022. Survey of the exocrine glands in workers of the ant Leptanilla clypeata. Insectes Sociaux -:1-13. 10.1007/s00040-022-00861 PDF

Mizuno, R.; Eguchi, K.; Satria, R.; Dang, A. V.; Bui, V. T.; Phung, L. T. H.; Ito, F. 2023. Colony composition, phasic reproduction, caste dimorphism, and prey preferences of the oriental non-army doryline ant Yunodorylus eguchii (Borowiec, 2009) (Hymenoptera: Formicidae: Dorylinae). Insectes Sociaux -:13 pp. [online early] 10.1007/s00040-023-00898-4 PDF

Ito, F.; Luong, P. T. H.; Yamane, S. 2023. Specialized predation on arthropod eggs in the myrmicine ant Calyptomyrmex rectopilosus collected in northern Vietnam, with a description of new species of Calyptomyrmex from Bogor, West Java, Indonesia. Tropics 32 (2):65-72. 10.3759/tropics.MS22-08 PDF

Yamane, S.; Ito, F. 2023. [Untitled. Calyptomyrmex nitidiusculus sp. nov. attributed to S. Yamane and F. Ito.]. Pp. 67-70 in: Ito, F.; Luong, P. T. H.; Yamane, S. 2023. Specialized predation on arthropod eggs in the myrmicine ant Calyptomyrmex rectopilosus collected in northern Vietnam, with a description of new species of Calyptomyrmex from Bogor, West Java, Indonesia. Tropics 32 (2):65-72. PDF

Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Erniwati; Kahono, S. 2024. Ant species diversity (Hymenoptera: Formicidae) in the Purwodadi Botanical Gardens, East Java, Indonesia. Treubia 51 (1):85-100. 10.14203/treubia.v51i1.4759 PDF

Khalife, A.; Sasaki, H.; Ito, F. 2024. Intercastes in the ant Aphaenogaster rugulosa (Formicidae: Myrmicinae): morphological variation and adaptive potential. Insectes Sociaux 71:11 pp. [online early] 10.1007/s00040-024-00987-y PDF

Sasaki, H.; Khalife, A.; Ito, F. 2025. Colony composition, prey preference and behavioral characteristics of Leptanilla kubotai (Hymenoptera: Formicidae: Leptanillinae). Asian Myrmecology 18:e018001, 9 pp. [online early] 10.20362/am.018001 PDF

Khachonpisitsak, S.; Jaitrong, W.; Jeenthong, T.; Wongkunanusorn, C.; Laedprathom, K.; Chaianunporn, T.; Khalife, A.; Aupanun, S.; Ito, F. 2025. Ant-plants as oases: Colonies of Cladomyrma sirindhornae (Hymenoptera, Formicidae) thrive in their host plants Sphenodesme pentandra in both the wet and dry seasons. Journal of Hymenoptera Research 98:559-577. [online early] 10.3897/jhr.98.144518 PDF