Wheeler, W. M. 1903i. The origin of female and worker ants from the eggs of parthenogenetic workers. Science (New York) (n.s.)18:830-833.

Wheeler, W. M. 1904b. Dr. Castle and the Dzierzon theory. Science (New York) (n.s.)19:587-591. PDF

Wheeler, W. M. 1904g. On the pupation of ants and the feasibility of establishing the Guatemalan Kelep, or Cotton-Weevil Ant, in the United States. Science (New York) (n.s.)20:437-440. PDF

Wheeler, W. M. 1904j. Some further comments on the Guatemalan boll weevil ant. Science (New York) (n.s.)20:766-768. PDF

Wheeler, W. M. 1905i. Dr. O. F. Cook's "Social organization and breeding habits of the cotton-protecting kelep of Guatemala.". Science (New York) (n.s.)21:706-710. PDF

Wheeler, W. M. 1906c. The kelep excused. Science (New York) (n.s.)23:348-350. PDF

Wheeler, W. M. 1911c. Three formicid names which have been overlooked. Science (New York) (n.s.)33:858-860. PDF

Wheeler, W. M. 1916e. [Review of: Donisthorpe, H. 1915. British ants, their life-history and classification. Plymouth: Brendon & Son Ltd., xv + 379 pp.]. Science (New York) (n.s.)43:316-318.

Wheeler, W. M. 1932f. How the primitive ants of Australia start their colonies. Science (New York) 76:532-533. PDF