Ward, P. S. 1989a. Systematic studies on pseudomyrmecine ants: revision of the Pseudomyrmex oculatus and P. subtilissimus species groups, with taxonomic comments on other species. Quaestiones Entomologicae 25:393-468. PDF

Ward, P. S. 1989b. Genetic and social changes associated with ant speciation. Pp. 123-148 in: Breed, M. D.; Page, R. E. (eds.) 1989. The genetics of social evolution. Boulder: Westview Press, 213 pp.

Ward, P. S. 1990. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): generic revision and relationship to other formicids. Systematic Entomology 15:449-489. PDF

Ward, P. S. 1991. Phylogenetic analysis of pseudomyrmecine ants associated with domatia-bearing plants. Pp. 335-352 in: Huxley, C. R.; Cutler, D. F. (eds.) 1991. Ant-plant interactions. Oxford: Oxford University Press, xviii + 601 pp. PDF

Ward, P. S. 1992. Ants of the genus Pseudomyrmex (Hymenoptera: Formicidae) from Dominican amber, with a synopsis of the extant Antillean species. Psyche (Cambridge) 99:55-85. PDF

Ward, P. S. 1993. Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae). Journal of Hymenoptera Research 2:117-168. PDF

Notes: Errata in J. Hym. Res. 3:309-310. (1994.10.15)

Ward, P. S. 1994. Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Systematic Entomology 19:159-175. PDF

Ward, P. S. 1995. Ants on the run. [Review of: Gotwald, W. H., Jr. 1995. Army ants: the biology of social predation. Ithaca, New York: Cornell University Press, xviii + 302 pp.]. Science (Washington, D. C.) 270:319-320. PDF

Ward, P. S. 1996. A new workerless social parasite in the ant genus Pseudomyrmex (Hymenoptera: Formicidae), with a discussion of the origin of social parasitism in ants. Systematic Entomology 21:253-263. PDF

Ward, P. S. 1997. Ant soldiers are not modified queens. Nature (London) 385:494-495. PDF

Ward, P. S. 1999a. Deceptive similarity in army ants of the genus Neivamyrmex (Hymenoptera: Formicidae): taxonomy, distribution and biology of N. californicus (Mayr) and N. nigrescens (Cresson). Journal of Hymenoptera Research 8:74-97. PDF

Ward, P. S. 1999b. Systematics, biogeography and host plant associations of the Pseudomyrmex viduus group (Hymenoptera: Formicidae), Triplaris- and Tachigali-inhabiting ants. Zoological Journal of the Linnean Society 126:451-540. PDF

Ward, P. S. 2000a. On the identity of Pheidole vaslitii Pergande (Hymenoptera: Formicidae), a neglected ant from Baja California. Journal of Hymenoptera Research 9:85-98. PDF

Ward, P. S. 2000b. Broad-scale patterns of diversity in leaf litter ant communities. Pp. 99-121 in: Agosti, D.; Majer, J. D.; Alonso, L. E.; Schultz, T. R. (eds.) 2000. Ants. Standard methods for measuring and monitoring biodiversity. Washington: Smithsonian Institution Press, xix + 280 pp. PDF

Ward, P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebrate Taxonomy 15:589-665. PDF

Ward, P. S. 2003. Subfamilia Pseudomyrmecinae. Pp. 331-333 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp. PDF

Ward, P. S. 2005. A synoptic review of the ants of California (Hymenoptera: Formicidae). Zootaxa 936:1-68. PDF

Ward, P. S. 2006. The ant genus Tetraponera in the Afrotropical region: synopsis of species groups and revision of the T. ambigua-group (Hymenoptera: Formicidae). Myrmecologische Nachrichten 8:119-130. PDF

Ward, P. S. 2007a. Edward O. Wilson and his contributions to ant systematics. Memoirs of the American Entomological Institute 80:3-7. PDF

Ward, P. S. 2007b. The ant genus Leptanilloides: discovery of the male and evaluation of phylogenetic relationships based on DNA sequence data. Pp. 637-649 in: Snelling, R. R.; Fisher, B. L.; Ward, P. S. (eds.) 2007. Advances in ant systematics (Hymenoptera: Formicidae): homage to E.O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute 80:690 pp. PDF

Ward, P. S. 2007c. Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae). Zootaxa 1668:549-563. PDF

Ward, P. S. 2009a. The ant genus Tetraponera in the Afrotropical region: the T. grandidieri group (Hymenoptera: Formicidae). Journal of Hymenoptera Research 18:285-304. PDF

Ward, P. S. 2009b. Taxonomy, phylogenetics, and evolution. Pp. 3-17 in: Lach, L.; Parr, C. L.; Abbott, K. (eds) 2009. Ant ecology. Oxford: Oxford University Press, xvii + 410 pp. PDF

Ward, P. S. 2011. Integrating molecular phylogenetic results into ant taxonomy (Hymenoptera: Formicidae). Myrmecological News 15:21-29. PDF

Ward, P. S. 2014. The phylogeny and evolution of ants. Annual Review of Ecology, Evolution, and Systematics 45:23-43. PDF

Ward, P. S. 2017. A review of the Pseudomyrmex ferrugineus and Pseudomyrmex goeldii species groups: acacia-ants and relatives (Hymenoptera: Formicidae). Zootaxa 4227 (4):524-542. 10.11646/zootaxa.4227.4.3 PDF

Ward, P. S. 2019. Subfamilia Pseudomyrmecinae. Capítulo 33. Pp. 1089-1113 in: Fernández, F.; Guerrero, R. J.; Delsinne, T. (eds.) 2019d. Hormigas de Colombia. Bogotá: Universidad Nacional de Colombia, 1198 pp. PDF

Ward, P. S. 2022. The ant genus Tetraponera (Hymenoptera: Formicidae) in the Afrotropical region: taxonomic review and key to species. Zootaxa 5102 (1):1-70. 10.11646/zootaxa.5102.1.1 PDF

Ward, P. S.; Blaimer, B. B. 2022. Taxonomy in the phylogenomic era: species boundaries and phylogenetic relationships among North American ants of the Crematogaster scutellaris group (Formicidae: Hymenoptera). Zoological Journal of the Linnean Society 194:893-937. PDF

Ward, P. S.; Blaimer, B. B.; Fisher, B. L. 2016. A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex. Zootaxa 4072:343-357. PDF