Ward, P. S. 1985b. The Nearctic species of the genus Pseudomyrmex (Hymenoptera: Formicidae). Quaestiones Entomologicae 21:209-246. PDF

Ward, P. S. 1986. Functional queens in the Australian greenhead ant, Rhytidoponera metallica (Hymenoptera: Formicidae). Psyche (Cambridge) 93:1-12. PDF

Ward, P. S. 1987. Distribution of the introduced Argentine ant (Iridomyrmex humilis) in natural habitats of the lower Sacramento Valley and its effects on the indigenous ant fauna. Hilgardia 55(2):1-16. PDF

Ward, P. S. 1988. Mesic elements in the western Nearctic ant fauna: taxonomic and biological notes on Amblyopone, Proceratium, and Smithistruma (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 61:102-124. PDF

Ward, P. S. 1989a. Systematic studies on pseudomyrmecine ants: revision of the Pseudomyrmex oculatus and P. subtilissimus species groups, with taxonomic comments on other species. Quaestiones Entomologicae 25:393-468. PDF

Ward, P. S. 1989b. Genetic and social changes associated with ant speciation. Pp. 123-148 in: Breed, M. D.; Page, R. E. (eds.) 1989. The genetics of social evolution. Boulder: Westview Press, 213 pp.

Ward, P. S. 1990. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): generic revision and relationship to other formicids. Systematic Entomology 15:449-489. PDF

Ward, P. S. 1991. Phylogenetic analysis of pseudomyrmecine ants associated with domatia-bearing plants. Pp. 335-352 in: Huxley, C. R.; Cutler, D. F. (eds.) 1991. Ant-plant interactions. Oxford: Oxford University Press, xviii + 601 pp. PDF

Ward, P. S. 1992. Ants of the genus Pseudomyrmex (Hymenoptera: Formicidae) from Dominican amber, with a synopsis of the extant Antillean species. Psyche (Cambridge) 99:55-85. PDF

Ward, P. S. 1993. Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae). Journal of Hymenoptera Research 2:117-168. PDF

Notes: Errata in J. Hym. Res. 3:309-310. (1994.10.15)

Ward, P. S. 1994. Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Systematic Entomology 19:159-175. PDF

Ward, P. S. 1995. Ants on the run. [Review of: Gotwald, W. H., Jr. 1995. Army ants: the biology of social predation. Ithaca, New York: Cornell University Press, xviii + 302 pp.]. Science (Washington, D. C.) 270:319-320. PDF

Ward, P. S. 1996. A new workerless social parasite in the ant genus Pseudomyrmex (Hymenoptera: Formicidae), with a discussion of the origin of social parasitism in ants. Systematic Entomology 21:253-263. PDF

Ward, P. S. 1997. Ant soldiers are not modified queens. Nature (London) 385:494-495. PDF

Ward, P. S. 1999a. Deceptive similarity in army ants of the genus Neivamyrmex (Hymenoptera: Formicidae): taxonomy, distribution and biology of N. californicus (Mayr) and N. nigrescens (Cresson). Journal of Hymenoptera Research 8:74-97. PDF

Ward, P. S. 1999b. Systematics, biogeography and host plant associations of the Pseudomyrmex viduus group (Hymenoptera: Formicidae), Triplaris- and Tachigali-inhabiting ants. Zoological Journal of the Linnean Society 126:451-540. PDF

Ward, P. S. 2000a. On the identity of Pheidole vaslitii Pergande (Hymenoptera: Formicidae), a neglected ant from Baja California. Journal of Hymenoptera Research 9:85-98. PDF

Ward, P. S. 2000b. Broad-scale patterns of diversity in leaf litter ant communities. Pp. 99-121 in: Agosti, D.; Majer, J. D.; Alonso, L. E.; Schultz, T. R. (eds.) 2000. Ants. Standard methods for measuring and monitoring biodiversity. Washington: Smithsonian Institution Press, xix + 280 pp. PDF

Ward, P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebrate Taxonomy 15:589-665. PDF

Ward, P. S. 2003. Subfamilia Pseudomyrmecinae. Pp. 331-333 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp. PDF

Ward, P. S. 2005. A synoptic review of the ants of California (Hymenoptera: Formicidae). Zootaxa 936:1-68. PDF

Ward, P. S. 2006. The ant genus Tetraponera in the Afrotropical region: synopsis of species groups and revision of the T. ambigua-group (Hymenoptera: Formicidae). Myrmecologische Nachrichten 8:119-130. PDF

Ward, P. S. 2007a. Edward O. Wilson and his contributions to ant systematics. Memoirs of the American Entomological Institute 80:3-7. PDF

Ward, P. S. 2007b. The ant genus Leptanilloides: discovery of the male and evaluation of phylogenetic relationships based on DNA sequence data. Pp. 637-649 in: Snelling, R. R.; Fisher, B. L.; Ward, P. S. (eds.) 2007. Advances in ant systematics (Hymenoptera: Formicidae): homage to E.O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute 80:690 pp. PDF

Ward, P. S. 2007c. Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae). Zootaxa 1668:549-563. PDF

Ward, P. S. 2009a. The ant genus Tetraponera in the Afrotropical region: the T. grandidieri group (Hymenoptera: Formicidae). Journal of Hymenoptera Research 18:285-304. PDF

Ward, P. S. 2009b. Taxonomy, phylogenetics, and evolution. Pp. 3-17 in: Lach, L.; Parr, C. L.; Abbott, K. (eds) 2009. Ant ecology. Oxford: Oxford University Press, xvii + 410 pp. PDF

Ward, P. S. 2011. Integrating molecular phylogenetic results into ant taxonomy (Hymenoptera: Formicidae). Myrmecological News 15:21-29. PDF

Ward, P. S. 2014. The phylogeny and evolution of ants. Annual Review of Ecology, Evolution, and Systematics 45:23-43. PDF

Ward, P. S. 2017. A review of the Pseudomyrmex ferrugineus and Pseudomyrmex goeldii species groups: acacia-ants and relatives (Hymenoptera: Formicidae). Zootaxa 4227 (4):524-542. 10.11646/zootaxa.4227.4.3 PDF