Ward, P. S. 1999b. Systematics, biogeography and host plant associations of the Pseudomyrmex viduus group (Hymenoptera: Formicidae), Triplaris- and Tachigali-inhabiting ants. Zoological Journal of the Linnean Society 126:451-540. PDF

Brady, S. G.; Gadau, J.; Ward, P. S. 2000. Systematics of the ant genus Camponotus (Hymenoptera: Formicidae): a preliminary analysis using data from the mitochondrial gene cytochrome oxidase I. Pp. 131-139 in: Austin, A. D.; Dowton, M. (eds.) 2000. Hymenoptera. Evolution, biodiversity and biological control. Collingwood, Victoria: CSIRO Publishing, xi + 468 pp.

Wetterer, J. K.; Ward, P. S.; Wetterer, A. L.; Longino, J. T.; Trager, J. C.; Miller, S. E. 2000. Ants (Hymenoptera: Formicidae) of Santa Cruz Island, California. Bulletin of the Southern California Academy of Sciences 99:25-31.

Ward, P. S. 2000a. On the identity of Pheidole vaslitii Pergande (Hymenoptera: Formicidae), a neglected ant from Baja California. Journal of Hymenoptera Research 9:85-98. PDF

Ward, P. S. 2000b. Broad-scale patterns of diversity in leaf litter ant communities. Pp. 99-121 in: Agosti, D.; Majer, J. D.; Alonso, L. E.; Schultz, T. R. (eds.) 2000. Ants. Standard methods for measuring and monitoring biodiversity. Washington: Smithsonian Institution Press, xix + 280 pp. PDF

Ward, P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebrate Taxonomy 15:589-665. PDF

Ward, P. S.; Brady, S. G. 2002. Phylogeny and biogeography of the ant genera Myrmecia and Nothomyrmecia (Hymenoptera: Formicidae) and related fossil taxa. [Abstract.]. P. 156 in: Billen, J. P. J. 2002. XIV International Congress of IUSSI. The Golden Jubilee Proceedings. 27 July - 3 August 2002, Hokkaido University, Sapporo, Japan. Sapporo: XIV International Congress of IUSSI, 233 pp.

Johnson, R. A.; Ward, P. S. 2002. Biogeography and endemism of ants (Hymenoptera: Formicidae) in Baja California, Mexico: a first overview. Journal of Biogeography 29:1009-1026. PDF

Ward, P. S.; Brady, S. G. 2003. Phylogeny and biogeography of the ant subfamily Myrmeciinae (Hymenoptera: Formicidae). Invertebrate Systematics 17:361-386. PDF

Ward, P. S. 2003. Subfamilia Pseudomyrmecinae. Pp. 331-333 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp. PDF

Kaspari, M.; Ward, P. S.; Yuan, M. 2004. Energy gradients and the geographic distribution of local ant diversity. Oecologia (Berlin) 140:407-413. PDF

Ward, P. S. 2005. A synoptic review of the ants of California (Hymenoptera: Formicidae). Zootaxa 936:1-68. PDF

Ward, P. S.; Downie, D. A. 2005. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): phylogeny and evolution of big-eyed arboreal ants. Systematic Entomology 30:310-335. PDF

Boulton, A. M.; Davies, K. F.; Ward, P. S. 2005. Species richness, abundance, and composition of ground-dwelling ants in northern California grasslands: role of plants, soil, and grazing. Environmental Entomology 34:96-104. PDF

Suarez, A. V.; Holway, D. A.; Ward, P. S. 2005. The role of opportunity in the unintentional introduction of nonnative ants. Proceedings of the National Academy of Sciences of the United States of America 102:17032-17035. PDF

van der Hammen, T.; Ward, P. S. 2005. Ants from the Ecoandes expeditions: diversity and distribution. Studies on Tropical Andean Ecosystems 6:239-248. PDF

Ward, P. S.; Brady, S. G.; Fisher, B. L.; Schultz, T. R. 2005. Assembling the ant "Tree of Life" (Hymenoptera: Formicidae). Myrmecologische Nachrichten 7:87-90. PDF

Brady, S. G.; Ward, P. S. 2005. Morphological phylogeny of army ants and other dorylomorphs (Hymenoptera: Formicidae). Systematic Entomology 30:593-618. PDF

Brady, S. G.; Schultz, T. R.; Fisher, B. L.; Ward, P. S. 2006. Evaluating alternative hypotheses for the early evolution and diversification of ants. Proceedings of the National Academy of Sciences of the United States of America 103:18172-18177. PDF

Ward, P. S. 2006. The ant genus Tetraponera in the Afrotropical region: synopsis of species groups and revision of the T. ambigua-group (Hymenoptera: Formicidae). Myrmecologische Nachrichten 8:119-130. PDF

Bolton, B.; Alpert, G.; Ward, P. S.; Naskrecki, P. 2007. Bolton's Catalogue of ants of the world: 1758-2005. Cambridge, Mass.: Harvard University Press, Compact Disk. PDF

Snelling, R. R.; Fisher, B. L.; Ward, P. S. (eds.) 2007. Advances in ant systematics (Hymenoptera: Formicidae): homage to E.O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute 80:690 pp.

Ward, P. S. 2007a. Edward O. Wilson and his contributions to ant systematics. Memoirs of the American Entomological Institute 80:3-7. PDF

Ward, P. S. 2007b. The ant genus Leptanilloides: discovery of the male and evaluation of phylogenetic relationships based on DNA sequence data. Pp. 637-649 in: Snelling, R. R.; Fisher, B. L.; Ward, P. S. (eds.) 2007. Advances in ant systematics (Hymenoptera: Formicidae): homage to E.O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute 80:690 pp. PDF

Ward, P. S. 2007c. Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae). Zootaxa 1668:549-563. PDF

Kautz, S.; Lumbsch, H. T.; Ward, P. S.; Heil, M. 2009. How to prevent cheating: a digestive specialization ties mutualistic plant-ants to their ant-plant partners. Evolution 63:839-853. PDF

Ward, P. S.; Brady, S. G. 2009. Rediscovery of the ant genus Amyrmex Kusnezov (Hymenoptera: Formicidae) and its transfer from Dolichoderinae to Leptanilloidinae. Zootaxa 2063:46-54. PDF

Wernegreen, J. J.; Kauppinen, S. N.; Brady, S. G.; Ward, P. S. 2009. One nutritional symbiosis begat another: phylogenetic evidence that the ant tribe Camponotini acquired Blochmannia by tending sap-feeding insects. BMC Evolutionary Biology 9:292. 10.1186/1471-2148-9-292 PDF

Ward, P. S. 2009a. The ant genus Tetraponera in the Afrotropical region: the T. grandidieri group (Hymenoptera: Formicidae). Journal of Hymenoptera Research 18:285-304. PDF

Ward, P. S. 2009b. Taxonomy, phylogenetics, and evolution. Pp. 3-17 in: Lach, L.; Parr, C. L.; Abbott, K. (eds) 2009. Ant ecology. Oxford: Oxford University Press, xvii + 410 pp. PDF