References by Ward, P. S.
This author has an antcat.org account
Ward, P. S. 1980a. A systematic revision of the Rhytidoponera impressa group (Hymenoptera: Formicidae) in Australia and New Guinea. Australian Journal of Zoology 28:475-498. PDF
Ward, P. S. 1980b. Genetic variation and population differentiation in the Rhytidoponera impressa group, a species complex of ponerine ants (Hymenoptera: Formicidae). Evolution 34:1060-1076. PDF
Ward, P. S.; Taylor, R. W. 1981. Allozyme variation, colony structure and genetic relatedness in the primitive ant Nothomyrmecia macrops Clark (Hymenoptera: Formicidae). Journal of the Australian Entomological Society 20:177-183. PDF
Ward, P. S. 1981a. Ecology and life history of the Rhytidoponera impressa group. I. Habitats, nest sites, and foraging behavior. Psyche (Cambridge) 88:89-108. PDF
Ward, P. S. 1981b. Ecology and life history of the Rhytidoponera impressa group. II. Colony origin, seasonal cycles, and reproduction. Psyche (Cambridge) 88:109-126. PDF
Ward, P. S. 1983a. Genetic relatedness and colony organization in a species complex of ponerine ants. I. Genotypic and phenotypic composition of colonies. Behavioral Ecology and Sociobiology 12:285-299. PDF
Ward, P. S. 1983b. Genetic relatedness and colony organization in a species complex of ponerine ants. II. Patterns of sex ratio investment. Behavioral Ecology and Sociobiology 12:301-307. PDF
Ward, P. S. 1984. A revision of the ant genus Rhytidoponera (Hymenoptera: Formicidae) in New Caledonia. Australian Journal of Zoology 32:131-175. PDF
Ward, P. S. 1985a. Taxonomic congruence and disparity in an insular ant fauna: Rhytidoponera in New Caledonia. Systematic Zoology 34:140-151. PDF
Ward, P. S. 1985b. The Nearctic species of the genus Pseudomyrmex (Hymenoptera: Formicidae). Quaestiones Entomologicae 21:209-246. PDF
Ward, P. S. 1986. Functional queens in the Australian greenhead ant, Rhytidoponera metallica (Hymenoptera: Formicidae). Psyche (Cambridge) 93:1-12. PDF
Ward, P. S. 1987. Distribution of the introduced Argentine ant (Iridomyrmex humilis) in natural habitats of the lower Sacramento Valley and its effects on the indigenous ant fauna. Hilgardia 55(2):1-16. PDF
Ward, P. S. 1988. Mesic elements in the western Nearctic ant fauna: taxonomic and biological notes on Amblyopone, Proceratium, and Smithistruma (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 61:102-124. PDF
Ward, P. S. 1989a. Systematic studies on pseudomyrmecine ants: revision of the Pseudomyrmex oculatus and P. subtilissimus species groups, with taxonomic comments on other species. Quaestiones Entomologicae 25:393-468. PDF
Ward, P. S. 1989b. Genetic and social changes associated with ant speciation. Pp. 123-148 in: Breed, M. D.; Page, R. E. (eds.) 1989. The genetics of social evolution. Boulder: Westview Press, 213 pp.
Ward, P. S. 1990. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): generic revision and relationship to other formicids. Systematic Entomology 15:449-489. PDF
Ward, P. S. 1991. Phylogenetic analysis of pseudomyrmecine ants associated with domatia-bearing plants. Pp. 335-352 in: Huxley, C. R.; Cutler, D. F. (eds.) 1991. Ant-plant interactions. Oxford: Oxford University Press, xviii + 601 pp. PDF
Baroni Urbani, C.; Bolton, B.; Ward, P. S. 1992. The internal phylogeny of ants (Hymenoptera: Formicidae). Systematic Entomology 17:301-329. PDF
Ward, P. S. 1992. Ants of the genus Pseudomyrmex (Hymenoptera: Formicidae) from Dominican amber, with a synopsis of the extant Antillean species. Psyche (Cambridge) 99:55-85. PDF
Frumhoff, P. C.; Ward, P. S. 1992. Individual-level selection, colony-level selection, and the association between polygyny and worker monomorphism in ants. American Naturalist 139:559-590. PDF
Ward, P. S. 1993. Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae). Journal of Hymenoptera Research 2:117-168. PDF

Notes: Errata in J. Hym. Res. 3:309-310. (1994.10.15)

Ward, P. S. 1994. Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Systematic Entomology 19:159-175. PDF
Ward, P. S. 1995. Ants on the run. [Review of: Gotwald, W. H., Jr. 1995. Army ants: the biology of social predation. Ithaca, New York: Cornell University Press, xviii + 302 pp.]. Science (Washington, D. C.) 270:319-320. PDF
Ward, P. S. 1996. A new workerless social parasite in the ant genus Pseudomyrmex (Hymenoptera: Formicidae), with a discussion of the origin of social parasitism in ants. Systematic Entomology 21:253-263. PDF
Ward, P. S.; Bolton, B.; Shattuck, S. O.; Brown, W. L., Jr. 1996. A bibliography of ant systematics. University of California Publications in Entomology 116:1-417. PDF
Olson, D. M.; Ward, P. S. 1996. The ant fauna (Hymenoptera: Formicidae) of Kirindy Forest (tropical dry forest) in western Madagascar. Pp. 161-164 in: Ganzhorn, J. U.; Sorg, J.-P. (eds.) 1996. Ecology and economy of a tropical dry forest in Madagascar. Primate Report 46-1. Göttingen: German Primate Center (DPZ), 382 pp. PDF
Ward, P. S. 1997. Ant soldiers are not modified queens. Nature (London) 385:494-495. PDF
Brady, S. G.; Gadau, J.; Ward, P. S. 1999. Is the ant genus Camponotus paraphyletic? [Abstract.]. Pp. 27-28 in: Publication Committee 1999. 4th International Hymenopterists Conference, 6-11th January 1999, Canberra, Australia. Program and abstracts. Glen Osmond, South Australia: 4th International Hymenopterists Conference, 96 pp.
Gadau, J.; Brady, S. G.; Ward, P. S. 1999. Systematics, distribution and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Annals of the Entomological Society of America 92:514-522. PDF
Ward, P. S. 1999a. Deceptive similarity in army ants of the genus Neivamyrmex (Hymenoptera: Formicidae): taxonomy, distribution and biology of N. californicus (Mayr) and N. nigrescens (Cresson). Journal of Hymenoptera Research 8:74-97. PDF
Protonyms originally described by author
Protonym Taxon Authorship Rank Status
Rhytidoponera confusa Rhytidoponera confusa Ward, 1980a Species Valid Original combination
Rhytidoponera enigmatica Rhytidoponera enigmatica Ward, 1980a Species Valid Original combination
Rhytidoponera aquila Rhytidoponera aquila Ward, 1984 Species Valid Original combination
Rhytidoponera arborea Rhytidoponera arborea Ward, 1984 Species Valid Original combination
Rhytidoponera depilis Rhytidoponera depilis Ward, 1984 Species Valid Original combination
Rhytidoponera insularis Rhytidoponera insularis Ward, 1984 Species Valid Original combination
Rhytidoponera koumensis Rhytidoponera koumensis Ward, 1984 Species Valid Original combination
Rhytidoponera litoralis Rhytidoponera litoralis Ward, 1984 Species Valid Original combination
Rhytidoponera luteipes Rhytidoponera luteipes Ward, 1984 Species Valid Original combination
Rhytidoponera mimica Rhytidoponera mimica Ward, 1984 Species Valid Original combination
Rhytidoponera nitidiventris Rhytidoponera nitidiventris Ward, 1984 Species Valid Original combination
Rhytidoponera opaciventris Rhytidoponera opaciventris Ward, 1984 Species Valid Original combination
Rhytidoponera terrestris Rhytidoponera terrestris Ward, 1984 Species Valid Original combination
Pseudomyrmex leptosus Pseudomyrmex leptosus Ward, 1985b Species Valid Original combination
Pseudomyrmex seminole Pseudomyrmex seminole Ward, 1985b Species Valid Original combination
Proceratium compitale Proceratium compitale Ward, 1988 Species Valid Original combination
Smithistruma arizonica Strumigenys arizonica Ward, 1988 Species Valid
Smithistruma chiricahua Strumigenys chiricahua Ward, 1988 Species Valid
Smithistruma reliquia Strumigenys reliquia Ward, 1988 Species Valid
Pseudomyrmex alustratus Pseudomyrmex alustratus Ward, 1989a Species Valid Original combination
Pseudomyrmex cretus Pseudomyrmex cretus Ward, 1989a Species Valid Original combination
Pseudomyrmex laevifrons Pseudomyrmex laevifrons Ward, 1989a Species Valid Original combination
Pseudomyrmex pisinnus Pseudomyrmex pisinnus Ward, 1989a Species Valid Original combination
Pseudomyrmex spiculus Pseudomyrmex spiculus Ward, 1989a Species Valid Original combination
Pseudomyrmex villosus Pseudomyrmex villosus Ward, 1989a Species Valid Original combination
Myrcidris Myrcidris Ward, 1990 Genus Valid
Myrcidris epicharis Myrcidris epicharis Ward, 1990 Species Valid Original combination
Leptanilloidinae Leptanilloidinae Baroni Urbani et al., 1992 Subfamily Synonym
†Pseudomyrmex antiquus †Pseudomyrmex antiquus Ward, 1992 Species Valid Original combination
†Pseudomyrmex avitus †Pseudomyrmex avitus Ward, 1992 Species Valid Original combination
Statistics
Published between 1980–2025
Taxon descriptions between 1980–2025
No. of references 89
No. of described taxa (including combinations) 127
No. of described taxa (valid, including combinations) 120
No. of described taxa (protonyms only) 123
Names belonging to author
Author name No. of references
Ward, P. S. 89