This history item belongs to Formica umbrata Nylander, 1846

Its current taxon owner(s) is Lasius umbratus

Status as species: Foerster, 1850a: 39; Schenck, 1852: 59; Mayr, 1855: 365 (redescription); Smith, 1855a: 106; Nylander, 1856b: 70; Gredler, 1858: 14; Smith, 1858a: 8, 52; Mayr, 1861: 50 (in key); Meinert, 1861: 320; Roger, 1863b: 11; Mayr, 1863a: 426; Emery, 1869b: 9; Smith, 1871c: 2; Dours, 1873: 165; Forel, 1874: 47 (in key); André, 1874b: 180 (in key); Emery, 1878a: ix (in list); Emery, 1878: 47; Emery & Forel, 1879: 452; Saunders, 1880: 209; André, 1882c: 195 (in key); White, 1884: 256; Mayr, 1886d: 430; Cresson, 1887: 257; Forel, 1889: 256; Nasonov, 1889: 24; Forel, 1892j: 307; Lameere, 1892: 64; Dalla Torre, 1893: 191; Ruzsky, 1896: 71; Saunders, 1896: 24; Forel, 1900h: 269; Forel, 1901m: 66; Ruzsky, 1902d: 14; Ruzsky, 1903c: 207; Ruzsky, 1905b: 284; Wheeler, 1906h: 322; Wasmann, 1906: 114 (in key); Bondroit, 1910: 485; Yano, 1910a: 421; Karavaiev, 1912b: 587; Ruzsky, 1914a: 61; Stitz, 1914: 86; Donisthorpe, 1915f: 223; Forel, 1915d: 52 (in key); Emery, 1916a: 241; Ruzsky, 1916: 5; Wheeler, 1916o: 172; Wheeler, 1917a: 528; Escherich, 1917: 333 (in key); Bondroit, 1918: 29; Nadig, 1918: 340; Bondroit, 1920a: 143; Emery, 1922b: 13; Finzi, 1922a: 120; Kulmatycki, 1922: 79; Soudek, 1922b: 72; Müller, 1923a: 75; Müller, 1923b: 129; Ruzsky, 1925b: 44; Emery, 1925d: 233; Karavaiev, 1926e: 195; Betrem, 1926: 215; Stärcke, 1926a: 122 (in key); Donisthorpe, 1927c: 262; Karavaiev, 1927a: 301; Karavaiev, 1927d: 279 (in key); Lomnicki, 1928: 7; Kuznetsov-Ugamsky, 1929b: 37; Karavaiev, 1930b: 148; Karavaiev, 1931e: 215; Soudek, 1931: 15; Arnol'di, 1933a: 603 (in key); Grandi, 1935: 103; Zimmermann, 1935: 52; Ruzsky, 1936: 91; Karavaiev, 1936: 211 (redescription); Stärcke, 1937: 39 (redescription); Stitz, 1939: 291; Teranishi, 1940: 22; Novák & Sadil, 1941: 103 (in key); Eidmann, 1941a: 26; Holgersen, 1943c: 174 (in key); Holgersen, 1944a: 181; Morisita, 1945: 22; Ruzsky, 1946: 70; Van Boven, 1947b: 185 (in key); Creighton, 1950a: 425; Azuma, 1951: 88; Chapman & Capco, 1951: 202; Azuma, 1955: 80; Wilson, 1955a: 150 (redescription); Ceballos, 1956: 316; Smith, 1958c: 150; Bernard, 1959a: 352; Carter, 1962a: 7 (in list); Collingwood, 1963b: 157 (in key); Bernard, 1967a: 363 (redescription); Smith, 1967a: 368; Kutter, 1968b: 61; Collingwood & Yarrow, 1969: 80; Baroni Urbani, 1971c: 213; Collingwood, 1971: 165; Banert & Pisarski, 1972: 354; Bourne, 1973: 25; Bolton & Collingwood, 1975: 7 (in key); Hunt & Snelling, 1975: 22; Pisarski, 1975: 38; Azuma, 1977a: 117; Van Boven, 1977: 151; Francoeur, 1977b: 207; Kutter, 1977c: 232; Yensen et al., 1977: 184; Collingwood, 1978: 90 (in key); Collingwood, 1979: 101; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 199; Allred, 1982: 483; Wheeler & Wheeler, 1986g: 67; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 281 (in key); Nilsson & Douwes, 1987: 70; Seifert, 1988a: 150 (redescription); DuBois & LaBerge, 1988: 149; MacKay et al., 1988: 118 (in key); Deyrup et al., 1989: 99; Casevitz-Weulersse, 1990c: 434; Seifert, 1990: 11; Morisita et al., 1991: 28; Atanassov & Dlussky, 1992: 245; Arakelian, 1994: 122; Radchenko, 1994b: 115 (in key); Wheeler et al., 1994: 308; Bolton, 1995b: 226; Douwes, 1995: 94; Poldi et al., 1995: 8; Espadaler, 1997g: 28; Gallé et al., 1998: 217; Czechowski et al., 2002: 110; Mackay & Mackay, 2002: 388; Deyrup, 2003: 45; Imai et al., 2003: 60; Coovert, 2005: 127; Csősz & Markó, 2005: 231; Karaman & Karaman, 2005: 58; MacGown & Forster, 2005: 64; Radchenko, 2005b: 172; Ward, 2005: 64; Bračko, 2006: 150; Markó et al., 2006: 69; Petrov, 2006: 106 (in key); Bračko, 2007: 20; Radchenko, 2007: 34; Seifert, 2007: 282; Werner & Wiezik, 2007: 142; Zryanin & Zryanina, 2007: 234; Gratiashvili & Barjadze, 2008: 135; Casevitz-Weulersse & Galkowski, 2009: 485; Lapeva-Gjonova et al., 2010: 41; Boer, 2010: 40; Csősz et al., 2011: 58; Karaman, 2011b: 93; Legakis, 2011: 27; Borowiec & Salata, 2012: 505; Czechowski et al., 2012: 274; Ellison et al., 2012: 203; Guénard & Dunn, 2012: 34; Kiran & Karaman, 2012: 13; Borowiec, 2014: 90; Bračko et al., 2014: 20; Lebas et al., 2016: 206; Radchenko, 2016: 373; Deyrup, 2017: 210; Steiner et al., 2017: 12; Salata & Borowiec, 2018c: 45; Schär et al., 2018: 6; Seifert, 2018: 285; Werner et al., 2018: 6; Borowiec & Salata, 2022: 222.


Status as species: {ref 124947}: 39; {ref 128471}: 59; {ref 127185}: 365 (redescription); {ref 128679}: 106; {ref 127565}: 70; {ref 125633}: 14; {ref 128685}: 8, 52; {ref 127189}: 50 (in key); {ref 127307}: 320; {ref 128094}: 11; {ref 127213}: 426; {ref 124512}: 9; {ref 128709}: 2; {ref 124387}: 165; {ref 124988}: 47 (in key); {ref 142475}: 180 (in key); {ref 124520}: ix (in list); {ref 132996}: 47; {ref 124778}: 452; {ref 128453}: 209; {ref 143315}: 195 (in key); {ref 130306}: 256; {ref 127240}: 430; {ref 123936}: 257; {ref 125018}: 256; {ref 127508}: 24; {ref 125040}: 307; {ref 126777}: 64; {ref 124002}: 191; {ref 128163}: 71; {ref 128456}: 24; {ref 125103}: 269; {ref 125117}: 66; {ref 128175}: 14; {ref 128180}: 207; {ref 128184}: 284; {ref 130013}: 322; {ref 129630}: 114 (in key); {ref 122874}: 485; {ref 130525}: 421; {ref 126240}: 587; {ref 128188}: 61; {ref 129025}: 86; {ref 124216}: 223; {ref 125260}: 52 (in key); {ref 124734}: 241; {ref 128165}: 5; {ref 130111}: 172; {ref 130116}: 528; {ref 124797}: 333 (in key); {ref 122872}: 29; {ref 127501}: 340; {ref 122879}: 143; {ref 124755}: 13; {ref 124918}: 120; {ref 126582}: 79; {ref 128940}: 72; {ref 127477}: 75; {ref 127478}: 129; {ref 128194}: 44; {ref 124775}: 233; {ref 126250}: 195; {ref 122702}: 215; {ref 143145}: 122 (in key); {ref 124250}: 262; {ref 126252}: 301; {ref 126255}: 279 (in key); {ref 126930}: 7; {ref 126763}: 37; {ref 126266}: 148; {ref 126271}: 215; {ref 128938}: 15; {ref 122356}: 603 (in key); {ref 125622}: 103; {ref 130613}: 52; {ref 128170}: 91; {ref 126235}: 211 (redescription); {ref 128991}: 39 (redescription); {ref 129033}: 291; {ref 129229}: 22; {ref 126512}: 103 (in key); {ref 124476}: 26; {ref 125920}: 174 (in key); {ref 125923}: 181; {ref 127446}: 22; {ref 128171}: 70; {ref 122970}: 185 (in key); {ref 123910}: 425; {ref 122419}: 88; {ref 123583}: 202; {ref 122421}: 80; {ref 130343}: 150 (redescription); {ref 123569}: 316; {ref 128854}: 150; {ref 122648}: 352; {ref 123542}: 7 (in list); {ref 123795}: 157 (in key); {ref 122660}: 363 (redescription); {ref 128863}: 368; {ref 126726}: 61; {ref 123816}: 80; {ref 122478}: 213; {ref 123761}: 165; {ref 122447}: 354; {ref 122950}: 25; {ref 122862}: 7 (in key); {ref 126032}: 22; {ref 127821}: 38; {ref 122424}: 117; {ref 122960}: 151; {ref 125337}: 207; {ref 126740}: 232; {ref 130568}: 184; {ref 123763}: 90 (in key); {ref 123764}: 101; {ref 128659}: 1439; {ref 130514}: 168; {ref 127629}: 199; {ref 122181}: 483; {ref 129925}: 67; {ref 122148}: 58; {ref 122149}: 281 (in key); {ref 127531}: 70; {ref 128568}: 150 (redescription); {ref 124419}: 149; {ref 127023}: 118 (in key); {ref 124103}: 99; {ref 123553}: 434; {ref 128560}: 11; {ref 127455}: 28; {ref 122405}: 245; {ref 122300}: 122; {ref 127961}: 115 (in key); {ref 129947}: 308; {ref 122860}: 226; {ref 124392}: 94; {ref 127868}: 8; {ref 131202}: 28; {ref 131353}: 217; {ref 130994}: 110; {ref 131795}: 388; {ref 131073}: 45; {ref 131519}: 60; {ref 143326}: 127; {ref 130956}: 231; {ref 132783}: 58; {ref 131768}: 64; {ref 132056}: 172; {ref 132562}: 64; {ref 132800}: 150; {ref 131816}: 69; {ref 132014}: 106 (in key); {ref 130818}: 20; {ref 132061}: 34; {ref 132271}: 282; {ref 132940}: 142; {ref 142997}: 234; {ref 131396}: 135; {ref 130895}: 485; {ref 132768}: 41; {ref 130781}: 40; {ref 132876}: 58; {ref 142634}: 93; {ref 132927}: 27; {ref 142313}: 505; {ref 142239}: 274; {ref 142320}: 203; {ref 142253}: 34; {ref 142245}: 13; {ref 142632}: 90; {ref 142696}: 20; {ref 143395}: 206; {ref 143043}: 373; {ref 143305}: 210; {ref 143816}: 12; {ref 143275}: 45; {ref 143185}: 6; {ref 143563}: 285; {ref 143817}: 6; {ref 144091}: 222.
  • Item type: taxt
  • Created at: 2012-09-12
  • Updated at: 2023-04-26
Related records
Taxa belonging to this history item's protonym
Formica umbrata Species Obsolete combination
an obsolete combination of Lasius umbratus (Nylander, 1846)
Formicina umbrata Species Obsolete combination
an obsolete combination of Lasius umbratus (Nylander, 1846)
Lasius umbratus Species Valid
Lasius umbratus umbratus Subspecies Obsolete combination
an obsolete combination of Lasius umbratus (Nylander, 1846)
Other history items belonging to Formica umbrata
#308930 Lectotype designation: Stärcke, 1937: 39.
#281499 [Misspelled as umbraus by Ruzsky, 1914a: 61.]
#257344 Schenck, 1852: 59 (w.); Hauschteck, 1962: 219 (k.); Hung, 1969: 456 (k.); Borowiec & Salata, 2022: 222 (w.)
#257345 Combination in Lasius: Mayr, 1861: 50 (in key).
#281500 Combination in Donisthorpea: Donisthorpe, 1915f: 223.
#281501 Combination in Lasius (Lasius): Forel, 1915d: 52.
#281502 Combination in Formicina: Emery, 1916b: 63; Bondroit, 1918: 29.
#281503 Combination in Formicina (Formicina): Emery, 1916a: 241.
#281504 Combination in Lasius (Formicina): Wheeler, 1916o: 172; Wheeler, 1917k: 463.
#281505 Combination in Acanthomyops: Ruzsky, 1925b: 44.
#281506 Combination in Acanthomyops (Chthonolasius): Betrem, 1926: 215.
#281507 Combination in Chthonolasius: Kulmatycki, 1922: 79; Ruzsky, 1936: 91.
#281508 Combination in Lasius (Chthonolasius): Ruzsky, 1912: 633; Ruzsky, 1914a: 59; Müller, 1923b: 129; Emery, 1925d: 233; Wilson, 1955a: 150.
#257347 (selected) Status as species: Foerster, 1850a: 39; Schenck, 1852: 59; Mayr, 1855: 365 (redescription); Smith, 1855a: 106; Nylander, 1856b: 70; Gredler, 1858: 14; Smith, 1858a: 8, 52; Mayr, 1861: 50 (in key); Meinert, 1861: 320; Roger, 1863b: 11; Mayr, 1863a: 426; Emery, 1869b: 9; Smith, 1871c: 2; Dours, 1873: 165; Forel, 1874: 47 (in key); André, 1874b: 180 (in key); Emery, 1878a: ix (in list); Emery, 1878: 47; Emery & Forel, 1879: 452; Saunders, 1880: 209; André, 1882c: 195 (in key); White, 1884: 256; Mayr, 1886d: 430; Cresson, 1887: 257; Forel, 1889: 256; Nasonov, 1889: 24; Forel, 1892j: 307; Lameere, 1892: 64; Dalla Torre, 1893: 191; Ruzsky, 1896: 71; Saunders, 1896: 24; Forel, 1900h: 269; Forel, 1901m: 66; Ruzsky, 1902d: 14; Ruzsky, 1903c: 207; Ruzsky, 1905b: 284; Wheeler, 1906h: 322; Wasmann, 1906: 114 (in key); Bondroit, 1910: 485; Yano, 1910a: 421; Karavaiev, 1912b: 587; Ruzsky, 1914a: 61; Stitz, 1914: 86; Donisthorpe, 1915f: 223; Forel, 1915d: 52 (in key); Emery, 1916a: 241; Ruzsky, 1916: 5; Wheeler, 1916o: 172; Wheeler, 1917a: 528; Escherich, 1917: 333 (in key); Bondroit, 1918: 29; Nadig, 1918: 340; Bondroit, 1920a: 143; Emery, 1922b: 13; Finzi, 1922a: 120; Kulmatycki, 1922: 79; Soudek, 1922b: 72; Müller, 1923a: 75; Müller, 1923b: 129; Ruzsky, 1925b: 44; Emery, 1925d: 233; Karavaiev, 1926e: 195; Betrem, 1926: 215; Stärcke, 1926a: 122 (in key); Donisthorpe, 1927c: 262; Karavaiev, 1927a: 301; Karavaiev, 1927d: 279 (in key); Lomnicki, 1928: 7; Kuznetsov-Ugamsky, 1929b: 37; Karavaiev, 1930b: 148; Karavaiev, 1931e: 215; Soudek, 1931: 15; Arnol'di, 1933a: 603 (in key); Grandi, 1935: 103; Zimmermann, 1935: 52; Ruzsky, 1936: 91; Karavaiev, 1936: 211 (redescription); Stärcke, 1937: 39 (redescription); Stitz, 1939: 291; Teranishi, 1940: 22; Novák & Sadil, 1941: 103 (in key); Eidmann, 1941a: 26; Holgersen, 1943c: 174 (in key); Holgersen, 1944a: 181; Morisita, 1945: 22; Ruzsky, 1946: 70; Van Boven, 1947b: 185 (in key); Creighton, 1950a: 425; Azuma, 1951: 88; Chapman & Capco, 1951: 202; Azuma, 1955: 80; Wilson, 1955a: 150 (redescription); Ceballos, 1956: 316; Smith, 1958c: 150; Bernard, 1959a: 352; Carter, 1962a: 7 (in list); Collingwood, 1963b: 157 (in key); Bernard, 1967a: 363 (redescription); Smith, 1967a: 368; Kutter, 1968b: 61; Collingwood & Yarrow, 1969: 80; Baroni Urbani, 1971c: 213; Collingwood, 1971: 165; Banert & Pisarski, 1972: 354; Bourne, 1973: 25; Bolton & Collingwood, 1975: 7 (in key); Hunt & Snelling, 1975: 22; Pisarski, 1975: 38; Azuma, 1977a: 117; Van Boven, 1977: 151; Francoeur, 1977b: 207; Kutter, 1977c: 232; Yensen et al., 1977: 184; Collingwood, 1978: 90 (in key); Collingwood, 1979: 101; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 199; Allred, 1982: 483; Wheeler & Wheeler, 1986g: 67; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 281 (in key); Nilsson & Douwes, 1987: 70; Seifert, 1988a: 150 (redescription); DuBois & LaBerge, 1988: 149; MacKay et al., 1988: 118 (in key); Deyrup et al., 1989: 99; Casevitz-Weulersse, 1990c: 434; Seifert, 1990: 11; Morisita et al., 1991: 28; Atanassov & Dlussky, 1992: 245; Arakelian, 1994: 122; Radchenko, 1994b: 115 (in key); Wheeler et al., 1994: 308; Bolton, 1995b: 226; Douwes, 1995: 94; Poldi et al., 1995: 8; Espadaler, 1997g: 28; Gallé et al., 1998: 217; Czechowski et al., 2002: 110; Mackay & Mackay, 2002: 388; Deyrup, 2003: 45; Imai et al., 2003: 60; Coovert, 2005: 127; Csősz & Markó, 2005: 231; Karaman & Karaman, 2005: 58; MacGown & Forster, 2005: 64; Radchenko, 2005b: 172; Ward, 2005: 64; Bračko, 2006: 150; Markó et al., 2006: 69; Petrov, 2006: 106 (in key); Bračko, 2007: 20; Radchenko, 2007: 34; Seifert, 2007: 282; Werner & Wiezik, 2007: 142; Zryanin & Zryanina, 2007: 234; Gratiashvili & Barjadze, 2008: 135; Casevitz-Weulersse & Galkowski, 2009: 485; Lapeva-Gjonova et al., 2010: 41; Boer, 2010: 40; Csősz et al., 2011: 58; Karaman, 2011b: 93; Legakis, 2011: 27; Borowiec & Salata, 2012: 505; Czechowski et al., 2012: 274; Ellison et al., 2012: 203; Guénard & Dunn, 2012: 34; Kiran & Karaman, 2012: 13; Borowiec, 2014: 90; Bračko et al., 2014: 20; Lebas et al., 2016: 206; Radchenko, 2016: 373; Deyrup, 2017: 210; Steiner et al., 2017: 12; Salata & Borowiec, 2018c: 45; Schär et al., 2018: 6; Seifert, 2018: 285; Werner et al., 2018: 6; Borowiec & Salata, 2022: 222.
#257346 Senior synonym of Lasius umbratus affinoumbratus: Wilson, 1955a: 150; Smith, 1979: 1439; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281509 Senior synonym of Lasius belgarum: Wilson, 1955a: 150; Bernard, 1967a: 363; Smith, 1979: 1439; Seifert, 1988a: 150; Seifert, 1990: 11; Bolton, 1995b: 226; Casevitz-Weulersse & Galkowski, 2009: 484; Radchenko, 2016: 373.
#281510 Senior synonym of Lasius exacutus: Wilson, 1955a: 150; Smith, 1979: 1439; Arakelian, 1994: 122; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281511 Senior synonym of Lasius umbratus hirtiscapus: Wilson, 1955a: 150; Smith, 1979: 1439; Seifert, 1990: 11; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281512 Senior synonym of Lasius affinis nyaradi: Wilson, 1955a: 150; Smith, 1979: 1439; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281513 Senior synonym of Lasius silvestrii osakana: Wilson, 1955a: 150; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 200; Bolton, 1995b: 226; Imai et al., 2003: 60; Radchenko, 2016: 373.
#281514 Senior synonym of Lasius silvestrii: Wilson, 1955a: 150; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 200; Bolton, 1995b: 226; Imai et al., 2003: 60; Radchenko, 2016: 373.
#281515 Material of the nomina nuda Lasius umbratus mixtoaffinis, Lasius umbratus mixtobicornis referred here by Wilson, 1955a: 150; Bolton, 1995b: 226.