This history item belongs to Formica umbrata Nylander, 1846

Its current taxon owner(s) is Lasius umbratus

Material of the nomina nuda Lasius umbratus mixtoaffinis, Lasius umbratus mixtobicornis referred here by Wilson, 1955a: 150; Bolton, 1995b: 226.


Material of the <i>nomina nuda</i> {tax 438817}, {tax 438818} referred here by {ref 130343}: 150; {ref 122860}: 226.
  • Item type: taxt
  • Created at: 2020-06-17
  • Updated at: 2020-06-17
Related records
Taxa belonging to this history item's protonym
Formica umbrata Species Obsolete combination
an obsolete combination of Lasius umbratus (Nylander, 1846)
Formicina umbrata Species Obsolete combination
an obsolete combination of Lasius umbratus (Nylander, 1846)
Lasius umbratus Species Valid
Lasius umbratus umbratus Subspecies Obsolete combination
an obsolete combination of Lasius umbratus (Nylander, 1846)
Other history items belonging to Formica umbrata
#308930 Lectotype designation: Stärcke, 1937: 39.
#281499 [Misspelled as umbraus by Ruzsky, 1914a: 61.]
#257344 Schenck, 1852: 59 (w.); Hauschteck, 1962: 219 (k.); Hung, 1969: 456 (k.); Borowiec & Salata, 2022: 222 (w.)
#257345 Combination in Lasius: Mayr, 1861: 50 (in key).
#281500 Combination in Donisthorpea: Donisthorpe, 1915f: 223.
#281501 Combination in Lasius (Lasius): Forel, 1915d: 52.
#281502 Combination in Formicina: Emery, 1916b: 63; Bondroit, 1918: 29.
#281503 Combination in Formicina (Formicina): Emery, 1916a: 241.
#281504 Combination in Lasius (Formicina): Wheeler, 1916o: 172; Wheeler, 1917k: 463.
#281505 Combination in Acanthomyops: Ruzsky, 1925b: 44.
#281506 Combination in Acanthomyops (Chthonolasius): Betrem, 1926: 215.
#281507 Combination in Chthonolasius: Kulmatycki, 1922: 79; Ruzsky, 1936: 91.
#281508 Combination in Lasius (Chthonolasius): Ruzsky, 1912: 633; Ruzsky, 1914a: 59; Müller, 1923b: 129; Emery, 1925d: 233; Wilson, 1955a: 150.
#257347 Status as species: Foerster, 1850a: 39; Schenck, 1852: 59; Mayr, 1855: 365 (redescription); Smith, 1855a: 106; Nylander, 1856b: 70; Gredler, 1858: 14; Smith, 1858a: 8, 52; Mayr, 1861: 50 (in key); Meinert, 1861: 320; Roger, 1863b: 11; Mayr, 1863a: 426; Emery, 1869b: 9; Smith, 1871c: 2; Dours, 1873: 165; Forel, 1874: 47 (in key); André, 1874b: 180 (in key); Emery, 1878a: ix (in list); Emery, 1878: 47; Emery & Forel, 1879: 452; Saunders, 1880: 209; André, 1882c: 195 (in key); White, 1884: 256; Mayr, 1886d: 430; Cresson, 1887: 257; Forel, 1889: 256; Nasonov, 1889: 24; Forel, 1892j: 307; Lameere, 1892: 64; Dalla Torre, 1893: 191; Ruzsky, 1896: 71; Saunders, 1896: 24; Forel, 1900h: 269; Forel, 1901m: 66; Ruzsky, 1902d: 14; Ruzsky, 1903c: 207; Ruzsky, 1905b: 284; Wheeler, 1906h: 322; Wasmann, 1906: 114 (in key); Bondroit, 1910: 485; Yano, 1910a: 421; Karavaiev, 1912b: 587; Ruzsky, 1914a: 61; Stitz, 1914: 86; Donisthorpe, 1915f: 223; Forel, 1915d: 52 (in key); Emery, 1916a: 241; Ruzsky, 1916: 5; Wheeler, 1916o: 172; Wheeler, 1917a: 528; Escherich, 1917: 333 (in key); Bondroit, 1918: 29; Nadig, 1918: 340; Bondroit, 1920a: 143; Emery, 1922b: 13; Finzi, 1922a: 120; Kulmatycki, 1922: 79; Soudek, 1922b: 72; Müller, 1923a: 75; Müller, 1923b: 129; Ruzsky, 1925b: 44; Emery, 1925d: 233; Karavaiev, 1926e: 195; Betrem, 1926: 215; Stärcke, 1926a: 122 (in key); Donisthorpe, 1927c: 262; Karavaiev, 1927a: 301; Karavaiev, 1927d: 279 (in key); Lomnicki, 1928: 7; Kuznetsov-Ugamsky, 1929b: 37; Karavaiev, 1930b: 148; Karavaiev, 1931e: 215; Soudek, 1931: 15; Arnol'di, 1933a: 603 (in key); Grandi, 1935: 103; Zimmermann, 1935: 52; Ruzsky, 1936: 91; Karavaiev, 1936: 211 (redescription); Stärcke, 1937: 39 (redescription); Stitz, 1939: 291; Teranishi, 1940: 22; Novák & Sadil, 1941: 103 (in key); Eidmann, 1941a: 26; Holgersen, 1943c: 174 (in key); Holgersen, 1944a: 181; Morisita, 1945: 22; Ruzsky, 1946: 70; Van Boven, 1947b: 185 (in key); Creighton, 1950a: 425; Azuma, 1951: 88; Chapman & Capco, 1951: 202; Azuma, 1955: 80; Wilson, 1955a: 150 (redescription); Ceballos, 1956: 316; Smith, 1958c: 150; Bernard, 1959a: 352; Carter, 1962a: 7 (in list); Collingwood, 1963b: 157 (in key); Bernard, 1967a: 363 (redescription); Smith, 1967a: 368; Kutter, 1968b: 61; Collingwood & Yarrow, 1969: 80; Baroni Urbani, 1971c: 213; Collingwood, 1971: 165; Banert & Pisarski, 1972: 354; Bourne, 1973: 25; Bolton & Collingwood, 1975: 7 (in key); Hunt & Snelling, 1975: 22; Pisarski, 1975: 38; Azuma, 1977a: 117; Van Boven, 1977: 151; Francoeur, 1977b: 207; Kutter, 1977c: 232; Yensen et al., 1977: 184; Collingwood, 1978: 90 (in key); Collingwood, 1979: 101; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 199; Allred, 1982: 483; Wheeler & Wheeler, 1986g: 67; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 281 (in key); Nilsson & Douwes, 1987: 70; Seifert, 1988a: 150 (redescription); DuBois & LaBerge, 1988: 149; MacKay et al., 1988: 118 (in key); Deyrup et al., 1989: 99; Casevitz-Weulersse, 1990c: 434; Seifert, 1990: 11; Morisita et al., 1991: 28; Atanassov & Dlussky, 1992: 245; Arakelian, 1994: 122; Radchenko, 1994b: 115 (in key); Wheeler et al., 1994: 308; Bolton, 1995b: 226; Douwes, 1995: 94; Poldi et al., 1995: 8; Espadaler, 1997g: 28; Gallé et al., 1998: 217; Czechowski et al., 2002: 110; Mackay & Mackay, 2002: 388; Deyrup, 2003: 45; Imai et al., 2003: 60; Coovert, 2005: 127; Csősz & Markó, 2005: 231; Karaman & Karaman, 2005: 58; MacGown & Forster, 2005: 64; Radchenko, 2005b: 172; Ward, 2005: 64; Bračko, 2006: 150; Markó et al., 2006: 69; Petrov, 2006: 106 (in key); Bračko, 2007: 20; Radchenko, 2007: 34; Seifert, 2007: 282; Werner & Wiezik, 2007: 142; Zryanin & Zryanina, 2007: 234; Gratiashvili & Barjadze, 2008: 135; Casevitz-Weulersse & Galkowski, 2009: 485; Lapeva-Gjonova et al., 2010: 41; Boer, 2010: 40; Csősz et al., 2011: 58; Karaman, 2011b: 93; Legakis, 2011: 27; Borowiec & Salata, 2012: 505; Czechowski et al., 2012: 274; Ellison et al., 2012: 203; Guénard & Dunn, 2012: 34; Kiran & Karaman, 2012: 13; Borowiec, 2014: 90; Bračko et al., 2014: 20; Lebas et al., 2016: 206; Radchenko, 2016: 373; Deyrup, 2017: 210; Steiner et al., 2017: 12; Salata & Borowiec, 2018c: 45; Schär et al., 2018: 6; Seifert, 2018: 285; Werner et al., 2018: 6; Borowiec & Salata, 2022: 222.
#257346 Senior synonym of Lasius umbratus affinoumbratus: Wilson, 1955a: 150; Smith, 1979: 1439; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281509 Senior synonym of Lasius belgarum: Wilson, 1955a: 150; Bernard, 1967a: 363; Smith, 1979: 1439; Seifert, 1988a: 150; Seifert, 1990: 11; Bolton, 1995b: 226; Casevitz-Weulersse & Galkowski, 2009: 484; Radchenko, 2016: 373.
#281510 Senior synonym of Lasius exacutus: Wilson, 1955a: 150; Smith, 1979: 1439; Arakelian, 1994: 122; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281511 Senior synonym of Lasius umbratus hirtiscapus: Wilson, 1955a: 150; Smith, 1979: 1439; Seifert, 1990: 11; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281512 Senior synonym of Lasius affinis nyaradi: Wilson, 1955a: 150; Smith, 1979: 1439; Bolton, 1995b: 226; Radchenko, 2016: 373.
#281513 Senior synonym of Lasius silvestrii osakana: Wilson, 1955a: 150; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 200; Bolton, 1995b: 226; Imai et al., 2003: 60; Radchenko, 2016: 373.
#281514 Senior synonym of Lasius silvestrii: Wilson, 1955a: 150; Smith, 1979: 1439; Yamauchi, 1979: 168; Onoyama, 1980a: 200; Bolton, 1995b: 226; Imai et al., 2003: 60; Radchenko, 2016: 373.
#281515 (selected) Material of the nomina nuda Lasius umbratus mixtoaffinis, Lasius umbratus mixtobicornis referred here by Wilson, 1955a: 150; Bolton, 1995b: 226.