This history item belongs to Formica aliena Foerster, 1850

Its current taxon owner(s) is Lasius alienus

Status as species: Schenck, 1852: 51; Mayr, 1855: 360 (redescription); Nylander, 1856b: 68; Gredler, 1858: 13; Smith, 1858a: 7; Mayr, 1861: 49 (in key); Roger, 1861b: 165; Roger, 1863b: 11; Mayr, 1863a: 425; Smith, 1871c: 2; Dours, 1873: 165; André, 1874b: 180 (in key); Mayr, 1877a: 6; Emery, 1878: 47; Saunders, 1880: 209; Mayr, 1880: 26; André, 1882c: 192 (in key); White, 1884: 255; Nasonov, 1889: 22; Emery, 1891c: 16; Dalla Torre, 1893: 181; Emery, 1897g: 238; Ruzsky, 1902d: 16; Ruzsky, 1903c: 207; Bingham, 1903: 342; Viehmeyer, 1906: 56; Emery, 1908e: 24; Bondroit, 1911a: 11; Donisthorpe, 1915f: 212; Bondroit, 1918: 25; Crawley, 1920b: 177; Müller, 1923a: 74; Müller, 1923b: 125; Lomnicki, 1925b: 3; Betrem, 1926: 215; Stärcke, 1926a: 124 (in key); Donisthorpe, 1927a: 8; Donisthorpe, 1927c: 242; Finzi, 1933: 165; Stitz, 1934: 9; Zimmermann, 1935: 48; Ruzsky, 1936: 90; Finzi, 1939c: 160; Stitz, 1939: 279; Novák & Sadil, 1941: 101 (in key); Röszler, 1942a: 53; Holgersen, 1943c: 174 (in key); Holgersen, 1944a: 180; Stärcke, 1944b: 153; Van Boven, 1947b: 186 (in key); Arnol'di, 1948a: 212 (in list); Creighton, 1950a: 419; Röszler, 1951: 91; Consani & Zangheri, 1952: 44; Wilson, 1955a: 77 (redescription); Bernard, 1956b: 261; Smith, 1958c: 147; Bernard, 1959a: 351; Carter, 1962a: 7 (in list); Dlussky, 1962: 182; Baroni Urbani, 1964a: 7; Baroni Urbani, 1964b: 63; Baroni Urbani, 1964c: 166; Cagniant, 1964: 92; Cagniant, 1966b: 281; Beck et al., 1967: 70; Bernard, 1967a: 356 (redescription); Smith, 1967a: 367; Baroni Urbani, 1968e: 488; Cagniant, 1968a: 146; Kutter, 1968b: 61; Baroni Urbani, 1969e: 334; Collingwood & Yarrow, 1969: 79; Pisarski, 1969a: 231; Pisarski, 1969b: 306; Dlussky & Pisarski, 1970: 87; Cagniant, 1970c: 38; Baroni Urbani, 1971c: 200; Collingwood, 1971: 165; Bourne, 1973: 24; Baroni Urbani, 1974a: 238; Bolton & Collingwood, 1975: 7 (in key); Hunt & Snelling, 1975: 22; Pisarski, 1975: 34; Tarbinsky, 1976: 138 (redescription); Collingwood, 1976a: 305; Aktaç, 1977: 126; Azuma, 1977a: 117; Van Boven, 1977: 144; Francoeur, 1977b: 207; Kutter, 1977c: 227; Yensen et al., 1977: 184; Arnol'di & Dlussky, 1978: 555 (in key); Collingwood, 1978: 89 (in key); Wheeler & Wheeler, 1978b: 393; Smith, 1979: 1435; Collingwood, 1979: 97; Yamauchi, 1979: 156; Onoyama, 1980a: 199; Collingwood, 1981: 27; Dlussky, 1981b: 17; Pisarski & Krzysztofiak, 1981: 160; Schembri & Collingwood, 1981: 438; Allred, 1982: 479; Collingwood, 1982: 285; Wheeler & Wheeler, 1986g: 65; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 282 (in key); Nilsson & Douwes, 1987: 70; DuBois & LaBerge, 1988: 149; Kugler, 1988: 259; MacKay et al., 1988: 118; Deyrup et al., 1989: 99; Kupyanskaya, 1990a: 218; Dlussky et al., 1990: 159; Casevitz-Weulersse, 1990c: 428; Le Moli & Rosi, 1991: 36; Morisita et al., 1991: 27; Atanassov & Dlussky, 1992: 237; Blacker, 1992: 9; Seifert, 1992b: 13 (redescription); Wu & Wang, 1992c: 1312; Collingwood, 1993b: 195; Arakelian, 1994: 116; Radchenko, 1994b: 115 (in key); Wheeler et al., 1994: 307; Bolton, 1995b: 221; Douwes, 1995: 94; Poldi et al., 1995: 7; Tang et al., 1995: 108; Wu & Wang, 1995a: 155; Espadaler, 1997g: 28; Collingwood & Prince, 1998: 23 (in key); Gallé et al., 1998: 216; Czechowski et al., 2002: 105; Mackay & Mackay, 2002: 379; Zhang & Zheng, 2002: 219; Deyrup, 2003: 45; Imai et al., 2003: 63; Karaman & Karaman, 2003: 52; Coovert, 2005: 120; Csősz & Markó, 2005: 230; Karaman & Karaman, 2005: 56; MacGown & Forster, 2005: 64; Ward, 2005: 64; Bračko, 2006: 148; Markó et al., 2006: 68; Petrov, 2006: 69, 107 (in key); Schultz et al., 2006: 205; Zhou, 2006: 587; Bračko, 2007: 20; Seifert, 2007: 272; Werner & Wiezik, 2007: 153; Zryanin & Zryanina, 2007: 234; Gratiashvili & Barjadze, 2008: 136; Paknia et al., 2008: 154; Borowiec et al., 2009: 378; Casevitz-Weulersse & Galkowski, 2009: 483; Lapeva-Gjonova et al., 2010: 35; Boer, 2010: 42; Csősz et al., 2011: 58; Karaman, 2011b: 86; Legakis, 2011: 26; Borowiec & Salata, 2012: 498; Czechowski et al., 2012: 263; Ellison et al., 2012: 189; Guénard & Dunn, 2012: 32; Kiran & Karaman, 2012: 11; Bharti & Gul, 2013a: 57 (in key); Borowiec, 2014: 83; Tohmé & Tohmé, 2014: 139; Bharti et al., 2016: 27; Lebas et al., 2016: 210; Radchenko, 2016: 362; Seifert & Galkowski, 2016: 52 (in key); Deyrup, 2017: 206; Steiner et al., 2017: 10; Borowiec & Salata, 2018a: 6; Salata & Borowiec, 2018c: 45; Schär et al., 2018: 6; Seifert, 2018: 268; Werner et al., 2018: 6; Rasheed et al., 2019: 430; Seifert, 2020a: 61; Arcos et al., 2022: 10; Borowiec & Salata, 2022: 186; Bračko, 2023: 150; Sert & Karaman, 2024: 397.


Status as species: {ref 128471}: 51; {ref 127185}: 360 (redescription); {ref 127565}: 68; {ref 125633}: 13; {ref 128685}: 7; {ref 127189}: 49 (in key); {ref 128089}: 165; {ref 128094}: 11; {ref 127213}: 425; {ref 128709}: 2; {ref 124387}: 165; {ref 142475}: 180 (in key); {ref 127231}: 6; {ref 132996}: 47; {ref 128453}: 209; {ref 127198}: 26; {ref 143315}: 192 (in key); {ref 130306}: 255; {ref 127508}: 22; {ref 124554}: 16; {ref 124002}: 181; {ref 124624}: 238; {ref 128175}: 16; {ref 128180}: 207; {ref 122766}: 342; {ref 129511}: 56; {ref 124680}: 24; {ref 122875}: 11; {ref 124216}: 212; {ref 122872}: 25; {ref 123869}: 177; {ref 127477}: 74; {ref 127478}: 125; {ref 126936}: 3; {ref 122702}: 215; {ref 143145}: 124 (in key); {ref 124248}: 8; {ref 124250}: 242; {ref 124916}: 165; {ref 129030}: 9; {ref 130613}: 48; {ref 128170}: 90; {ref 124938}: 160; {ref 129033}: 279; {ref 126512}: 101 (in key); {ref 128152}: 53; {ref 125920}: 174 (in key); {ref 125923}: 180; {ref 129007}: 153; {ref 122970}: 186 (in key); {ref 122358}: 212 (in list); {ref 123910}: 419; {ref 128144}: 91; {ref 123832}: 44; {ref 130343}: 77 (redescription); {ref 122644}: 261; {ref 128854}: 147; {ref 122648}: 351; {ref 123542}: 7 (in list); {ref 124124}: 182; {ref 122461}: 7; {ref 122462}: 63; {ref 122463}: 166; {ref 123447}: 92; {ref 123449}: 281; {ref 122564}: 70; {ref 122660}: 356 (redescription); {ref 128863}: 367; {ref 122468}: 488; {ref 123455}: 146; {ref 126726}: 61; {ref 122474}: 334; {ref 123816}: 79; {ref 127828}: 231; {ref 127829}: 306; {ref 124166}: 87; {ref 123464}: 38; {ref 122478}: 200; {ref 123761}: 165; {ref 122950}: 24; {ref 122482}: 238; {ref 122862}: 7 (in key); {ref 126032}: 22; {ref 127821}: 34; {ref 129149}: 138 (redescription); {ref 123801}: 305; {ref 122158}: 126; {ref 122424}: 117; {ref 122960}: 144; {ref 125337}: 207; {ref 126740}: 227; {ref 130568}: 184; {ref 122371}: 555 (in key); {ref 123763}: 89 (in key); {ref 129910}: 393; {ref 128659}: 1435; {ref 123764}: 97; {ref 130514}: 156; {ref 127629}: 199; {ref 123765}: 27; {ref 124147}: 17; {ref 127839}: 160; {ref 128469}: 438; {ref 122181}: 479; {ref 123766}: 285; {ref 129925}: 65; {ref 122148}: 58; {ref 122149}: 282 (in key); {ref 127531}: 70; {ref 124419}: 149; {ref 126577}: 259; {ref 127023}: 118; {ref 124103}: 99; {ref 126604}: 218; {ref 124151}: 159; {ref 123553}: 428; {ref 126846}: 36; {ref 127455}: 27; {ref 122405}: 237; {ref 122776}: 9; {ref 128573}: 13 (redescription); {ref 130486}: 1312; {ref 123807}: 195; {ref 122300}: 116; {ref 127961}: 115 (in key); {ref 129947}: 307; {ref 122860}: 221; {ref 124392}: 94; {ref 127868}: 7; {ref 129145}: 108; {ref 130487}: 155; {ref 131202}: 28; {ref 130938}: 23 (in key); {ref 131353}: 216; {ref 130994}: 105; {ref 131795}: 379; {ref 132705}: 219; {ref 131073}: 45; {ref 131519}: 63; {ref 132782}: 52; {ref 143326}: 120; {ref 130956}: 230; {ref 132783}: 56; {ref 131768}: 64; {ref 132562}: 64; {ref 132800}: 148; {ref 131816}: 68; {ref 132014}: 69, 107 (in key); {ref 132242}: 205; {ref 143253}: 587; {ref 130818}: 20; {ref 132271}: 272; {ref 132940}: 153; {ref 142997}: 234; {ref 131396}: 136; {ref 143565}: 154; {ref 130813}: 378; {ref 130895}: 483; {ref 132768}: 35; {ref 130781}: 42; {ref 132876}: 58; {ref 142634}: 86; {ref 132927}: 26; {ref 142313}: 498; {ref 142239}: 263; {ref 142320}: 189; {ref 142253}: 32; {ref 142245}: 11; {ref 142339}: 57 (in key); {ref 142632}: 83; {ref 143644}: 139; {ref 142873}: 27; {ref 143395}: 210; {ref 143043}: 362; {ref 142947}: 52 (in key); {ref 143305}: 206; {ref 143816}: 10; {ref 143244}: 6; {ref 143275}: 45; {ref 143185}: 6; {ref 143563}: 268; {ref 143817}: 6; {ref 143716}: 430; {ref 143711}: 61; {ref 144063}: 10; {ref 144091}: 186; {ref 144399}: 150; {ref 144400}: 397.
  • Item type: taxt
  • Created at: 2012-09-12
  • Updated at: 2024-09-10
Related records
Taxa belonging to this history item's protonym
Acanthomyops niger alienus Subspecies Obsolete combination
an obsolete combination of Lasius alienus (Foerster, 1850)
Formica aliena Species Obsolete combination
an obsolete combination of Lasius alienus (Foerster, 1850)
Lasius alienus Species Valid
Lasius niger alienus Subspecies Obsolete combination
an obsolete combination of Lasius alienus (Foerster, 1850)
Other history items belonging to Formica aliena
#308783 Neotype designation: Seifert, 1992b: 13 (invalid).
#308782 Neotype designation: Seifert, 2020a: 61.
#297426 [Misspelled as alineus by Pisarski & Krzysztofiak, 1981: 160.]
#256934 Foerster, 1850a: 71 (q.); Wheeler & Wheeler, 1953c: 147 (l.); Hauschteck, 1962: 219 (k.)
#256935 Combination in Lasius: Mayr, 1861: 49 (in key).
#297419 Combination in Lasius (Lasius): Ruzsky, 1912: 633; Forel, 1915d: 53; Wheeler, 1916o: 172; Karavaiev, 1936: 205.
#286069 Combination in Donisthorpea: Donisthorpe, 1915f: 212.
#286068 Combination in Formicina (Donisthorpea): Emery, 1916a: 240.
#297420 Combination in Formicina: Bondroit, 1918: 25.
#297421 Combination in Lasius (Donisthorpea): Nadig, 1918: 340.
#286067 Combination in Acanthomyops: Ruzsky, 1925b: 44.
#297422 Combination in Acanthomyops (Donisthorpea): Betrem, 1926: 215; Donisthorpe, 1927a: 8; Donisthorpe, 1950e: 1063.
#286066 Combination in Lasius: Ruzsky, 1916: 5; Wheeler, 1917k: 463; Menozzi, 1921: 32; Müller, 1923b: 125; Emery, 1925d: 230; Kuznetsov-Ugamsky, 1929a: 27.
#297423 Combination in Lasius (Lasius): Wilson, 1955a: 77; Seifert, 2020a: 16.
#256936 Subspecies of Lasius niger: Forel, 1874: 46 (in key); Mayr, 1877a: 20 (in list); Emery & Forel, 1879: 452; Mayr, 1886d: 429; Forel, 1886e: clxvii; Provancher, 1887: 236 (in key); Cresson, 1887: 255; Forel, 1889: 256; Forel, 1890b: lxvii; Emery, 1891c: 16; Forel, 1892j: 307; Lameere, 1892: 64; Emery, 1893m: 187 (footnote); Forel, 1894c: 404; Forel, 1894d: 12; Ruzsky, 1896: 71; Saunders, 1896: 25; Ruzsky, 1902a: 233; Ruzsky, 1903b: 306; Forel, 1904c: 386; Ruzsky, 1905b: 304; Wheeler, 1906h: 322; Forel, 1906b: 85; Forel, 1906c: 189; Wasmann, 1906: 114 (in key); Forel, 1909c: 105; Yano, 1910a: 421; Bondroit, 1910: 485; Forel, 1910a: 23; Karavaiev, 1910: 268; Forel, 1911f: 352; Karavaiev, 1912b: 588; Krausse, 1912c: 165; Forel, 1913d: 438; Ruzsky, 1914a: 61; Stitz, 1914: 84; Emery, 1914c: 159; Ruzsky, 1915a: 434; Forel, 1915d: 53 (in key); Emery, 1916a: 240; Ruzsky, 1916: 5; Wheeler, 1916o: 172; Escherich, 1917: 332 (in key); Wheeler, 1917a: 525; Wheeler, 1917k: 463; Nadig, 1918: 340; Santschi, 1919e: 246; Menozzi, 1921: 32; Menozzi, 1922c: 331; Soudek, 1922b: 69; Kulmatycki, 1922: 80; Finzi, 1923a: 4; Kuznetsov-Ugamsky, 1923b: 243; Emery, 1925d: 230; Ruzsky, 1925a: 288; Ruzsky, 1925b: 44; Santschi, 1925g: 349; Schkaff, 1925b: 275; Soudek, 1925b: 16; Santschi, 1926f: 289; Menozzi, 1926b: 182; Karavaiev, 1927a: 300; Karavaiev, 1927d: 280 (in key); Karavaiev, 1927e: 347; Kuznetsov-Ugamsky, 1927e: 188; Menozzi, 1927b: 91; Kuznetsov-Ugamsky, 1928b: 20; Lomnicki, 1928: 8; Wheeler, 1928c: 38; Kuznetsov-Ugamsky, 1929a: 27; Kuznetsov-Ugamsky, 1929b: 37; Karavaiev, 1930b: 147; Wheeler, 1930k: 80; Karavaiev, 1931e: 214; Soudek, 1931: 13; Gösswald, 1932: 57; Menozzi, 1932b: 311; Wheeler, 1932a: 16; Arnol'di, 1933a: 603 (in key); Grandi, 1935: 103; Karavaiev, 1935b: 108; Menozzi, 1936b: 305; Karavaiev, 1936: 205 (redescription); Karavaiev, 1937: 175; Menozzi, 1939a: 312; Morisita, 1945: 22; Ruzsky, 1946: 69; Azuma, 1951: 88; Smith, 1951c: 850; Chapman & Capco, 1951: 202; Azuma, 1953: 4; Azuma, 1955: 80; Ceballos, 1956: 316.
#256937 (selected) Status as species: Schenck, 1852: 51; Mayr, 1855: 360 (redescription); Nylander, 1856b: 68; Gredler, 1858: 13; Smith, 1858a: 7; Mayr, 1861: 49 (in key); Roger, 1861b: 165; Roger, 1863b: 11; Mayr, 1863a: 425; Smith, 1871c: 2; Dours, 1873: 165; André, 1874b: 180 (in key); Mayr, 1877a: 6; Emery, 1878: 47; Saunders, 1880: 209; Mayr, 1880: 26; André, 1882c: 192 (in key); White, 1884: 255; Nasonov, 1889: 22; Emery, 1891c: 16; Dalla Torre, 1893: 181; Emery, 1897g: 238; Ruzsky, 1902d: 16; Ruzsky, 1903c: 207; Bingham, 1903: 342; Viehmeyer, 1906: 56; Emery, 1908e: 24; Bondroit, 1911a: 11; Donisthorpe, 1915f: 212; Bondroit, 1918: 25; Crawley, 1920b: 177; Müller, 1923a: 74; Müller, 1923b: 125; Lomnicki, 1925b: 3; Betrem, 1926: 215; Stärcke, 1926a: 124 (in key); Donisthorpe, 1927a: 8; Donisthorpe, 1927c: 242; Finzi, 1933: 165; Stitz, 1934: 9; Zimmermann, 1935: 48; Ruzsky, 1936: 90; Finzi, 1939c: 160; Stitz, 1939: 279; Novák & Sadil, 1941: 101 (in key); Röszler, 1942a: 53; Holgersen, 1943c: 174 (in key); Holgersen, 1944a: 180; Stärcke, 1944b: 153; Van Boven, 1947b: 186 (in key); Arnol'di, 1948a: 212 (in list); Creighton, 1950a: 419; Röszler, 1951: 91; Consani & Zangheri, 1952: 44; Wilson, 1955a: 77 (redescription); Bernard, 1956b: 261; Smith, 1958c: 147; Bernard, 1959a: 351; Carter, 1962a: 7 (in list); Dlussky, 1962: 182; Baroni Urbani, 1964a: 7; Baroni Urbani, 1964b: 63; Baroni Urbani, 1964c: 166; Cagniant, 1964: 92; Cagniant, 1966b: 281; Beck et al., 1967: 70; Bernard, 1967a: 356 (redescription); Smith, 1967a: 367; Baroni Urbani, 1968e: 488; Cagniant, 1968a: 146; Kutter, 1968b: 61; Baroni Urbani, 1969e: 334; Collingwood & Yarrow, 1969: 79; Pisarski, 1969a: 231; Pisarski, 1969b: 306; Dlussky & Pisarski, 1970: 87; Cagniant, 1970c: 38; Baroni Urbani, 1971c: 200; Collingwood, 1971: 165; Bourne, 1973: 24; Baroni Urbani, 1974a: 238; Bolton & Collingwood, 1975: 7 (in key); Hunt & Snelling, 1975: 22; Pisarski, 1975: 34; Tarbinsky, 1976: 138 (redescription); Collingwood, 1976a: 305; Aktaç, 1977: 126; Azuma, 1977a: 117; Van Boven, 1977: 144; Francoeur, 1977b: 207; Kutter, 1977c: 227; Yensen et al., 1977: 184; Arnol'di & Dlussky, 1978: 555 (in key); Collingwood, 1978: 89 (in key); Wheeler & Wheeler, 1978b: 393; Smith, 1979: 1435; Collingwood, 1979: 97; Yamauchi, 1979: 156; Onoyama, 1980a: 199; Collingwood, 1981: 27; Dlussky, 1981b: 17; Pisarski & Krzysztofiak, 1981: 160; Schembri & Collingwood, 1981: 438; Allred, 1982: 479; Collingwood, 1982: 285; Wheeler & Wheeler, 1986g: 65; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 282 (in key); Nilsson & Douwes, 1987: 70; DuBois & LaBerge, 1988: 149; Kugler, 1988: 259; MacKay et al., 1988: 118; Deyrup et al., 1989: 99; Kupyanskaya, 1990a: 218; Dlussky et al., 1990: 159; Casevitz-Weulersse, 1990c: 428; Le Moli & Rosi, 1991: 36; Morisita et al., 1991: 27; Atanassov & Dlussky, 1992: 237; Blacker, 1992: 9; Seifert, 1992b: 13 (redescription); Wu & Wang, 1992c: 1312; Collingwood, 1993b: 195; Arakelian, 1994: 116; Radchenko, 1994b: 115 (in key); Wheeler et al., 1994: 307; Bolton, 1995b: 221; Douwes, 1995: 94; Poldi et al., 1995: 7; Tang et al., 1995: 108; Wu & Wang, 1995a: 155; Espadaler, 1997g: 28; Collingwood & Prince, 1998: 23 (in key); Gallé et al., 1998: 216; Czechowski et al., 2002: 105; Mackay & Mackay, 2002: 379; Zhang & Zheng, 2002: 219; Deyrup, 2003: 45; Imai et al., 2003: 63; Karaman & Karaman, 2003: 52; Coovert, 2005: 120; Csősz & Markó, 2005: 230; Karaman & Karaman, 2005: 56; MacGown & Forster, 2005: 64; Ward, 2005: 64; Bračko, 2006: 148; Markó et al., 2006: 68; Petrov, 2006: 69, 107 (in key); Schultz et al., 2006: 205; Zhou, 2006: 587; Bračko, 2007: 20; Seifert, 2007: 272; Werner & Wiezik, 2007: 153; Zryanin & Zryanina, 2007: 234; Gratiashvili & Barjadze, 2008: 136; Paknia et al., 2008: 154; Borowiec et al., 2009: 378; Casevitz-Weulersse & Galkowski, 2009: 483; Lapeva-Gjonova et al., 2010: 35; Boer, 2010: 42; Csősz et al., 2011: 58; Karaman, 2011b: 86; Legakis, 2011: 26; Borowiec & Salata, 2012: 498; Czechowski et al., 2012: 263; Ellison et al., 2012: 189; Guénard & Dunn, 2012: 32; Kiran & Karaman, 2012: 11; Bharti & Gul, 2013a: 57 (in key); Borowiec, 2014: 83; Tohmé & Tohmé, 2014: 139; Bharti et al., 2016: 27; Lebas et al., 2016: 210; Radchenko, 2016: 362; Seifert & Galkowski, 2016: 52 (in key); Deyrup, 2017: 206; Steiner et al., 2017: 10; Borowiec & Salata, 2018a: 6; Salata & Borowiec, 2018c: 45; Schär et al., 2018: 6; Seifert, 2018: 268; Werner et al., 2018: 6; Rasheed et al., 2019: 430; Seifert, 2020a: 61; Arcos et al., 2022: 10; Borowiec & Salata, 2022: 186; Bračko, 2023: 150; Sert & Karaman, 2024: 397.
#256938 Material of the unavailable name Lasius niger alienus alienoamericanus referred here by Wilson, 1955a: 77; Smith, 1958c: 147; Smith, 1979: 1435.
#297424 Material of the unavailable names Lasius niger alienus flavidus, Lasius niger alienus turkmenus referred here by Wilson, 1955a: 77; Smith, 1979: 1435; Dlussky et al., 1990: 159.
#297425 Material of the unavailable name Lasius niger alienus pilicornis referred to Lasius niger by Wilson, 1955a: 59; Smith, 1979: 1436; referred to Lasius alienus by Dlussky et al., 1990: 159.