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An Online Catalog of the Ants of the World
by Barry Bolton
Catalog
›
Formicidae
›
Formicinae
›
Plagiolepidini
›
Lepisiota
›
Lepisiota frauenfeldi
Lepisiota
frauenfeldi
(Mayr, 1855)
valid
Hypoclinea frauenfeldi
Mayr, 1855
: 378 (w.) CROATIA. Palearctic.
Primary type information:
Primary type material: syntype workers (number not stated). Primary type locality: Croatia: Dalmatia, Sign (
G.R. von Frauenfeld
). Primary type depository:
NHMW
.
Roger, 1859
: 243 (q.);
Emery, 1878
: 46 (m.);
André, 1881c
: 63 (m.);
Athias-Henriot, 1947
: 249 (l.);
Wheeler & Wheeler, 1968a
: 209 (l.).
Combination in
Formica (Hypoclinea)
:
Roger, 1859
: 243.
Combination in
Acantholepis
:
Mayr, 1861
: 42 (in key).
Combination in
Lepisiota
:
Baroni Urbani
et al.
, 1992
: 303.
Status as species:
Smith, 1858a
: 57;
Roger, 1859
: 243;
Mayr, 1861
: 42;
Mayr, 1863a
: 394;
Roger, 1863b
: 11;
André, 1874b
: 179 (in key);
Mayr, 1877a
: 11;
Emery, 1878
: 46;
Emery & Forel, 1879
: 453;
Mayr, 1880
: 30;
Emery, 1880
: 389;
Emery, 1881b
: 527;
André, 1881c
: 61;
André, 1882c
: 210 (in key);
André, 1884f
: 537;
Forel, 1885b
: 177;
Forel, 1886e
: clxvii;
Forel, 1889
: 256;
Nasonov, 1889
: 25;
Forel, 1890b
: lxviii;
Emery, 1891c
: 16;
Forel, 1892a
: 41 (in key);
Dalla Torre, 1893
: 171;
Forel, 1894c
: 413;
Forel, 1894d
: 12;
Forel, 1895e
: 233;
Forel, 1902a
: 155;
Ruzsky, 1902d
: 18;
Ruzsky, 1903b
: 308;
Rothney, 1903
: 99;
Bingham, 1903
: 316;
Forel, 1904c
: 387;
Forel, 1904d
: 12;
Ruzsky, 1905b
: 459;
Karavaiev, 1911a
: 41;
Forel, 1910a
: 23;
Forel, 1911f
: 351;
Karavaiev, 1912a
: 15;
Karavaiev, 1912b
: 586;
Emery, 1914c
: 158;
Emery, 1916a
: 221;
Wheeler & Mann, 1916
: 172;
Santschi, 1917d
: 42;
Donisthorpe, 1918c
: 167;
Crawley, 1920b
: 177;
Müller, 1923b
: 115;
Emery, 1925d
: 25;
Soudek, 1925b
: 16;
Donisthorpe, 1927a
: 8;
Finzi, 1928c
: 791;
Kuznetsov-Ugamsky, 1929b
: 41;
Finzi, 1930c
: 23;
Finzi, 1930d
: 316;
Mukerjee, 1930
: 156;
Finzi, 1932a
: 24;
Karavaiev, 1932
: 250;
Grandi, 1935
: 102;
Zimmermann, 1935
: 44;
Finzi, 1936
: 184;
Finzi, 1940
: 162;
Athias-Henriot, 1947
: 249;
Arnol'di, 1948a
: 212 (in list);
Bernard, 1948
: 155;
Donisthorpe, 1950e
: 1063;
Weber, 1952c
: 20;
Bernard, 1956b
: 260;
Ceballos, 1956
: 310;
Collingwood, 1961a
: 72;
Collingwood, 1961c
: 289;
Bernard & Cagniant, 1963
: 162;
Baroni Urbani, 1964a
: 7;
Cagniant, 1966b
: 280;
Baroni Urbani, 1968e
: 480;
Cagniant, 1968a
: 145;
Cagniant, 1970c
: 30;
Baroni Urbani, 1971c
: 173;
Bernard, 1971
: 8;
Pisarski, 1975
: 28;
Tohmé & Tohmé, 1976
: 131 (redescription);
Tarbinsky, 1976
: 130;
Baroni Urbani, 1976b
: 215;
Aktaç, 1977
: 124;
Collingwood, 1978
: 88 (in key);
Schembri & Collingwood, 1981
: 436;
Tohmé, 1981
: 1;
Collingwood, 1985
: 295;
Agosti & Collingwood, 1987a
: 57;
Agosti & Collingwood, 1987b
: 281 (in key);
Atanassov & Dlussky, 1992
: 205;
Collingwood, 1993b
: 195;
Arakelian, 1994
: 80;
Bolton, 1995b
: 227;
Mei, 1995
: 767;
Poldi
et al.
, 1995
: 7;
Espadaler, 1997g
: 28;
Collingwood & Prince, 1998
: 22;
Tiwari, 1999
: 67;
Mathew & Tiwari, 2000
: 347;
Blard
et al.
, 2003
: 129;
Bračko, 2006
: 150;
Petrov, 2006
: 105 (in key);
Gratiashvili & Barjadze, 2008
: 137;
Vonshak & Ionescu-Hirsch, 2010
: 40;
Lapeva-Gjonova
et al.
, 2010
: 34;
Karaman, 2011b
: 94;
Legakis, 2011
: 24;
Borowiec & Salata, 2012
: 506;
Kiran & Karaman, 2012
: 14;
Borowiec, 2014
: 93 (see note in bibliography);
Bračko
et al.
, 2014
: 20;
Tohmé & Tohmé, 2014
: 137;
Bharti
et al.
, 2016
: 28;
Lebas
et al.
, 2016
: 226;
Sharaf
et al.
, 2016
: 10 (in key) ;
Borowiec & Salata, 2018a
: 6;
Salata & Borowiec, 2018c
: 46;
Rasheed
et al.
, 2019
: 430;
Borowiec & Salata, 2020
: 10;
Sharaf
et al.
, 2020
: 133 (in key);
Dias
et al.
, 2020
: 42;
Arcos
et al.
, 2022
: 11;
Borowiec & Salata, 2022
: 228 (species complex);
Sert & Karaman, 2024
: 397.
Senior synonym of
Lepisiota frauenfeldi azerbeidzhanica
:
Arakelian, 1994
: 81;
Bolton, 1995b
: 227.
Junior synonyms
Lepisiota frauenfeldi azerbeidzhanica
(Karavaiev, 1932)
Obsolete combination(s):
Acantholepis frauenfeldi azerbeidzhanica
Obsolete combinations
Acantholepis frauenfeldi
Hypoclinea frauenfeldi
Current subspecies (nominal plus)
L. f. aegyptiaca
(Finzi, 1936)
L. f. atlantis
(Santschi, 1917)
L. f. barbara
(Santschi, 1917)
L. f. ferganica
(Kuznetsov-Ugamsky, 1929)
L. f. kantarensis
(Forel, 1911)
L. f. libanica
(Santschi, 1921)
L. f. marocana
(Santschi, 1936)
L. f. pubescens
(Forel, 1892)
L. f. saharensis
(Santschi, 1917)
L. f. truncata
(Finzi, 1940)
L. f. variabilis
(Santschi, 1917)
Citations
Agosti, D.
;
Collingwood, C. A.
1987a.
A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list.
Mitteilungen der Schweizerischen Entomologischen Gesellschaft 60:51-62.
PDF
Agosti, D.
;
Collingwood, C. A.
1987b.
A provisional list of the Balkan ants (Hym. Formicidae) with a key to the worker caste. II. Key to the worker caste, including the European species without the Iberian.
Mitteilungen der Schweizerischen Entomologischen Gesellschaft 60:261-293.
PDF
Aktaç, N.
1977 ("1976").
Studies on the myrmecofauna of Turkey I. Ants of Siirt, Bodrum and Trabzon.
Istanbul Universitesi Fen Fakultesi Mecmuasi. Seri B 41:115-135.
PDF
André, Ern.
1874b.
Description des fourmis d'Europe pour servir à l'étude des insectes myrmécophiles. [part].
Revue et Magasin de Zoologie Pure et Appliquée (3)2:161-192.
André, Ern.
1881c.
Catalogue raisonné des Formicides provenant du voyage en Orient de M. Abeille de Perrin et description des espèces nouvelles.
Annales de la Société Entomologique de France (6)1:53-78.
PDF
André, Ern.
1882c.
Les fourmis [part].
Pp. 153-232 in:
André, Edm.
1882.
1881-1886. Species des Hyménoptères d'Europe et d'Algérie. Tome Deuxième.
Beaune: Edmond André, 919 + 48 pp.
PDF
André, Ern.
1884f.
[Untitled.
Meranoplus Magrettii
, n. sp.].
P. 543 in:
Magretti, P.
1884.
Risultati di raccolte imenotterologiche nell'Africa Orientale.
Annali del Museo Civico di Storia Naturale 21:523-636.
Arakelian, G. R.
1994.
Fauna of the Republic of Armenia. Hymenopterous insects. Ants (Formicidae). [In Russian.].
Erevan: Gitutium, 153 pp.
PDF
Arcos, J.
;
Chaves, D.
;
Alarcón, P.
;
Rosado, Á.
2022.
First record of
Temnothorax convexus
(Forel, 1894) in Portugal (Hymenoptera: Formicidae) with an updated checklist of the ants from the country.
Sociobiology 69 (2): e7623:1-15.
10.13102/sociobiology.v69i2.7623
PDF
Arnol'di, K. V.
1948a.
Ants of Talysh and the Diabar depression. Their importance for the characterization of communities of terrestrial invertebrates and for historical analysis of the fauna. [In Russian.].
Trudy Zoologicheskogo Instituta. Akademiya Nauk SSSR 7(2):206-262.
PDF
Atanassov, N.
;
Dlussky, G. M.
1992.
Fauna of Bulgaria. Hymenoptera, Formicidae. [In Bulgarian.].
Fauna na Bûlgariya 22:1-310.
Athias-Henriot, C.
1947.
Recherches sur les larves de quelques fourmis D'Algérie.
Bulletin Biologique de la France et de la Belgique 81:247-272.
PDF
Baroni Urbani, C.
1964a.
Su alcune formiche raccolte in Turchia.
Annuario dell'Istituto e Museo di Zoologia dell'Università di Napoli 16:1-12.
PDF
Baroni Urbani, C.
1968e.
Studi sulla mirmecofauna d'Italia. IV. La fauna mirmecologica delle isole Maltesi ed il suo significato ecologico e biogeografico.
Annali del Museo Civico di Storia Naturale "Giacomo Doria" 77:408-559.
PDF
Baroni Urbani, C.
1971c.
Catalogo delle specie di Formicidae d'Italia (Studi sulla mirmecofauna d'Italia X).
Memorie della Società Entomologica Italiana 50:5-287.
PDF
Baroni Urbani, C.
1976b.
Le formiche dell'arcipelago della Galita (Tunisia).
Redia 59:207-223.
PDF
Baroni Urbani, C.
;
Bolton, B.
;
Ward, P. S.
1992.
The internal phylogeny of ants (Hymenoptera: Formicidae).
Systematic Entomology 17:301-329.
PDF
Bernard, F.
1948.
Les insectes sociaux du Fezzân. Comportement et biogéographie.
Pp. 87-200 in:
Bernard, F.
;
Peyerimhoff, P. de
1948.
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PDF
Bernard, F.
1956b.
Remarques sur le peuplement des Baléares en fourmis.
Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 47:254-266.
PDF
Bernard, F.
1971.
Les fourmis de l'Île de Djerba (Tunisie).
Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 62:3-14.
Bernard, F.
;
Cagniant, H.
1963 ("1962").
Capture au Hoggar de trois
Acantholepis
nouveaux pour ce massif avec observations sur leurs modes de vie (Hym. Formicidae).
Bulletin de la Société Entomologique de France 67:161-164.
PDF
Bharti, H.
;
Guénard, B.
;
Bharti, M.
;
Economo, E. P.
2016.
An updated checklist of the ants of India with their specific distributions in Indian states (Hymenoptera, Formicidae).
ZooKeys 551:1-83.
PDF
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PDF
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;
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;
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2003.
Les formis de l'île de la Réunion (Hymenoptera, Formicidae).
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PDF
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A new general catalogue of the ants of the world.
Cambridge, Mass.: Harvard University Press, 504 pp.
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2014.
Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae).
Genus (Wroclaw) 25 (1-2):1-340.
PDF
Borowiec, L.
;
Salata, S.
2012.
Ants of Greece – checklist, comments and new faunistic data (Hymenoptera: Formicidae).
Genus (Wroclaw) 23 (4):461-563.
PDF
Borowiec, L.
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2018a.
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10.5281/zenodo.1481802
PDF
Borowiec, L.
;
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2020.
Review of ants (Hymenoptera: Formicidae) from Jordan.
Annals of the Upper Silesian Museum, Entomology 29(002):1-29.
10.5281/zenodo.3733156
PDF
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;
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A monographic review of ants of Greece (Hymenoptera: Formicidae). Vol. 1. Introduction and review of all subfamilies except the subfamily Myrmicinae. Part 1: text.
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PDF
Bračko, G.
2006.
Review of the ant fauna (Hymenoptera: Formicidae) of Croatia.
Acta Entomologica Slovenica 14 (2):131-156.
PDF
Bračko, G.
;
Wagner, H. C.
;
Schulz, A.
;
Gioahin, E.
;
Matičič, J.
;
Tratnik, A.
2014.
New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia.
North-Western Journal of Zoology 10 (1):10-24.
PDF
Cagniant, H.
1966b.
Note sur le peuplement en fourmis d'une montagne de la région d'Alger, l'Atlas de Blida.
Bulletin de la Société d'Histoire Naturelle de Toulouse 102:278-284.
PDF
Cagniant, H.
1968a.
Liste préliminaire de fourmis forestières d'Algérie. Résultats obtenus de 1963 à 1966.
Bulletin de la Société d'Histoire Naturelle de Toulouse 104:138-147.
PDF
Cagniant, H.
1970c.
Deuxième liste de fourmis d'Algérie, récoltées principalement en forêt (Deuxième partie).
Bulletin de la Société d'Histoire Naturelle de Toulouse 106:28-40.
PDF
Ceballos, G.
1956.
Catálogo de los Himenópteros de España.
Madrid: Instituto Español de Entomología, 554 pp.
Collingwood, C. A.
1961a ("1960").
The third Danish Expedition to Central Asia. Zoological Results 27. Formicidae (Insecta) from Afghanistan.
Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 123:51-79.
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1961c.
Ergebnisse der Deutschen Afghanistan-Expedition 1956 der Landessammlungen für Naturkunde Karlsruhe. Formicidae (Hymenoptera Aculeata).
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1978.
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EOS. Revista Española de Entomología 52:65-95.
PDF
Collingwood, C. A.
1985.
Hymenoptera: Fam. Formicidae of Saudi Arabia.
Fauna of Saudi Arabia 7:230-302.
PDF
Collingwood, C. A.
1993b.
A comparative study of the ant fauna of five Greek islands.
Biologia Gallo-Hellenica 20:191-197.
PDF
Collingwood, C.
;
Prince, A.
1998.
A guide to ants of continental Portugal (Hymenoptera: Formicidae).
Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5:1-49.
Crawley, W. C.
1920b.
Ants from Mesopotamia and north-west Persia (concluded).
Entomologist's Record and Journal of Variation 32:177-179.
PDF
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1893.
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PDF
Dias, R. K. S.
;
Guénard, B.
;
Akbar, S. A.
;
Economo, E. P.
;
Udayakantha, W. S.
;
Wachkoo, A. A.
2020.
The ants (
Hymenoptera
,
Formicidae
) of Sri Lanka: a taxonomic research summary and updated checklist.
ZooKeys 967:1-142.
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PDF
Donisthorpe, H.
1918c.
A list of ants from Mesopotamia; with a description of a new species and a new variety.
Entomologist's Record and Journal of Variation 30:165-167.
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Donisthorpe, H.
1927a.
The ants (Formicidae), and some myrmecophiles, of Sicily. [concl.].
Entomologist's Record and Journal of Variation 39:6-9.
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Donisthorpe, H.
1950e.
A first instalment of the ants of Turkey.
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Emery, C.
1878.
Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte seconda. Formiche dell'Europa e delle regioni limitrofe in Africa e in Asia. Dummy reference.
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Emery, C.
1880.
Crociera del Violante, comandato dal capitano armatore Enrico d'Albertis, durante l'anno 1877. Formiche.
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PDF
Emery, C.
1881b.
Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso "Esploratore" dal 16 novembre 1879 al 26 febbraio 1880. I. Formiche.
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PDF
Emery, C.
1891c.
Exploration scientifique de la Tunisie. Zoologie. - Hyménoptères. Révision critique des fourmis de la Tunisie.
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Emery, C.
1914c.
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Emery, C.
1916a ("1915").
Fauna entomologica italiana. I. Hymenoptera.-Formicidae.
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Emery, C.
1925d.
Hymenoptera. Fam. Formicidae. Subfam. Formicinae.
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Emery, C.
;
Forel, A.
1879.
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PDF
Espadaler, X.
1997g.
Catàleg de les formigues (Hymenoptera: Formicidae) dels països catalans.
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PDF
Finzi, B.
1928c.
Weitere Beiträge zur Kenntnis der Fauna Griechenlands und der Inseln des Aegäischen Meeres. 1. Ameisen aus Griechenland und von den Aegäischen Inseln.
Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I 137:787-792.
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Finzi, B.
1930c.
Hymenopteren aus Palästina und Syrien. (Zoologische Studienreise von R. Ebner 1928 mit Unterstützung der Akademie der Wissenschaften in Wien.) Formicidae.
Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I 139:22-24.
PDF
Finzi, B.
1930d.
Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes. XII. Teil. Die Ameisen der Jonischen Inseln.
Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I 139:309-319.
PDF
Finzi, B.
1932a.
Sopra alcune formiche dell'Isola di Rodi.
Bollettino della Società Entomologica Italiana 64:24-27.
PDF
Finzi, B.
1936.
Risultati scientifici della spedizione di S. A. S. il Principe Alessandro della Torre e Tasso nell'Egitto e peninsola del Sinai. XI. Formiche.
Bulletin. Société Entomologique d'Egypte 20:155-210.
PDF
Finzi, B.
1940 ("1939").
Formiche della Libia.
Memorie della Società Entomologica Italiana 18:155-166.
PDF
Forel, A.
1885b.
Indian ants of the Indian Museum, Calcutta.
Journal of the Asiatic Society of Bengal. Part II. Natural Science 54:176-182.
PDF
Forel, A.
1886e.
Nouvelles fourmis de Grèce récoltées par M. E. von Oertzen.
Annales de la Société Entomologique de Belgique 30:clix-clxviii.
PDF
Forel, A.
1889 ("1888").
Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen.
Berliner Entomologische Zeitschrift 32:255-265.
PDF
Forel, A.
1890b.
Fourmis de Tunisie et de l'Algérie orientale.
Annales de la Société Entomologique de Belgique 34:lxi-lxxvi.
PDF
Forel, A.
1892a.
Notes myrmécologiques.
Annales de la Société Entomologique de Belgique 36:38-43.
PDF
Forel, A.
1894c.
Les Formicides de l'Empire des Indes et de Ceylan. Part IV.
Journal of the Bombay Natural History Society 8:396-420.
PDF
Forel, A.
1894d.
Les Formicides de la Province d'Oran (Algérie).
Bulletin de la Société Vaudoise des Sciences Naturelles 30:1-45.
PDF
Forel, A.
1895e.
Südpalaearctische Ameisen.
Mitteilungen der Schweizerischen Entomologischen Gesellschaft 9:227-234.
PDF
Forel, A.
1902a.
Les fourmis du Sahara algérien récoltées par M. le Professeur A. Lameere et le Dr. A. Diehl.
Annales de la Société Entomologique de Belgique 46:147-158.
PDF
Forel, A.
1904c ("1903").
Note sur les fourmis du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg.
Ezhegodnik Zoologicheskago Muzeya 8:368-388.
PDF
Forel, A.
1904d.
Miscellanea myrmécologiques.
Revue Suisse de Zoologie 12:1-52.
PDF
Forel, A.
1910a.
Glanures myrmécologiques.
Annales de la Société Entomologique de Belgique 54:6-32.
PDF
Forel, A.
1911f.
Fourmis nouvelles ou intéressantes.
Bulletin de la Société Vaudoise des Sciences Naturelles 47:331-400.
PDF
Grandi, G.
1935.
Contributi alla conoscenza degli Imenotteri Aculeati. XV.
Bollettino [del Reale Istituto] dell'Istituto di Entomologia della Università degli Studi di Bologna 8:27-121.
PDF
Gratiashvili, N.
;
Barjadze, S.
2008.
Checklist of the ants (Formicidae Latreille, 1809) of Georgia.
Proceedings of the Institute of Zoology (Tbilisi) 23:130-146.
PDF
Karaman, M. G.
2011b.
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Verhandlungen der Zoologisch-Botanischen Vereins in Wien 5:273-478.
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Journal of the Bombay Natural History Society 34:149-163.
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Le formiche della Venezia Guilia e della Dalmazia.
Bollettino della Società Adriatica di Scienze Naturali in Trieste 28:11-180.
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Extant: 11 valid subspecies
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obsolete combination
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acuta
affinis
ajjer
albata
alexis
ambigua
angolensis
annandalei
arabica
arenaria
arnoldi
aurea
bahrainensis
bicolor
binghami
bipartita
cacozela
canescens
capensis
capitata
carbonaria
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chapmani
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crinita
curta
dammama
dendrophila
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depressa
dhofara
dolabellae
egregia
elbazi
elegantissima
elevata
emmelii
erythraea
fergusoni
foreli
frauenfeldi
gerardi
gracilicornis
harteni
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integra
integrisquama
kabulica
karawaiewi
layla
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longinoda
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mayri
megacephala
melanogaster
melas
mlanjiensis
modesta
monardi
ngangela
nigra
nigrescens
nigrisetosa
nigriventris
obtusa
oculata
omanensis
opaca
opaciventris
palpalis
piliscapa
pulchella
pusaensis
quadraticeps
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riyadha
rothneyi
rubrovaria
rugithorax
satpuraensis
schoutedeni
semenovi
sericea
siamensis
silvicola
simplex
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spinisquama
spinosior
submetallica
surchanica
syriaca
tenuipilis
thepthepae
validiuscula
velox
wilsoni
wroughtonii
xichangensis
aegyptiaca
atlantis
barbara
ferganica
kantarensis
libanica
marocana
pubescens
saharensis
truncata
variabilis