Primary type information:
Primary type material: holotype queen. Primary type locality: Enspel Oilshale, Rhineland Palatinate, Germany. Enspel Formation, Upper Oligocene, MP 28; (24.56–24.79 Ma, Mertz
et al. 2007), layer S16. Primary type depository:
MNHM. Primary type specimen: NHMM-PE2001/5066-LS.